Precursor (bird)
Encyclopedia
"Precursor" is a controversial prehistoric bird
Bird
Birds are feathered, winged, bipedal, endothermic , egg-laying, vertebrate animals. Around 10,000 living species and 188 families makes them the most speciose class of tetrapod vertebrates. They inhabit ecosystems across the globe, from the Arctic to the Antarctic. Extant birds range in size from...

 genus
Genus
In biology, a genus is a low-level taxonomic rank used in the biological classification of living and fossil organisms, which is an example of definition by genus and differentia...

 from the Early Eocene
Eocene
The Eocene Epoch, lasting from about 56 to 34 million years ago , is a major division of the geologic timescale and the second epoch of the Paleogene Period in the Cenozoic Era. The Eocene spans the time from the end of the Palaeocene Epoch to the beginning of the Oligocene Epoch. The start of the...

. It was established based on fossil
Fossil
Fossils are the preserved remains or traces of animals , plants, and other organisms from the remote past...

s found in England
England
England is a country that is part of the United Kingdom. It shares land borders with Scotland to the north and Wales to the west; the Irish Sea is to the north west, the Celtic Sea to the south west, with the North Sea to the east and the English Channel to the south separating it from continental...

, e.g. in the famous London Clay
London Clay
The London Clay Formation is a marine geological formation of Ypresian age which crops out in the southeast of England. The London Clay is well known for the fossils it contains. The fossils from the Lower Eocene indicate a moderately warm climate, the flora being tropical or subtropical...

 deposits. Three species
Species
In biology, a species is one of the basic units of biological classification and a taxonomic rank. A species is often defined as a group of organisms capable of interbreeding and producing fertile offspring. While in many cases this definition is adequate, more precise or differing measures are...

 have been named, but the supposed "genus" seems to be a chimera
Chimera (paleontology)
In paleontology, a chimera is a fossil which was reconstructed with elements coming from more than a single species of animal. A now classic example of chimera is Protoavis.-List of paleontological chimeras:*Brontosaurus*Lametasaurus...

 composed from remains of different and quite unrelated Neornithes.

At least some of the remains seem to belong to a sort of psittaciform, of which today various parrot
Parrot
Parrots, also known as psittacines , are birds of the roughly 372 species in 86 genera that make up the order Psittaciformes, found in most tropical and subtropical regions. The order is subdivided into three families: the Psittacidae , the Cacatuidae and the Strigopidae...

s and cockatoo
Cockatoo
A cockatoo is any of the 21 species belonging to the bird family Cacatuidae. Along with the Psittacidae and the Strigopidae , they make up the parrot order Psittaciformes . Placement of the cockatoos as a separate family is fairly undisputed, although many aspects of the other living lineages of...

s exist. Initially, it was even supposed that the fossils belonged to some family
Family (biology)
In biological classification, family is* a taxonomic rank. Other well-known ranks are life, domain, kingdom, phylum, class, order, genus, and species, with family fitting between order and genus. As for the other well-known ranks, there is the option of an immediately lower rank, indicated by the...

 that is still extant. By now, it seems more likely however that the psittaciform material of the "Precursor" paratype
Paratype
Paratype is a technical term used in the scientific naming of species and other taxa of organisms. The exact meaning of the term paratype when it is used in zoology is not the same as the meaning when it is used in botany...

 belongs into the Pseudasturidae (of which Halcyornithidae may be the correct name). These were either basal in or immediately outside the Psittaciformes, and looked more like puffbird
Puffbird
The puffbirds and their relatives in the near passerine family Bucconidae are tropical birds breeding from South America up to Mexico.They are related to the jacamars, but lack the iridescent colours of that family. They are mainly brown, rufous or grey, with large heads and flattened bills with a...

s in habitus
Morphology (biology)
In biology, morphology is a branch of bioscience dealing with the study of the form and structure of organisms and their specific structural features....

 though they were almost certainly not at all closely related to the latter. True parrots probably originated in the approximate region of Australia
Australia
Australia , officially the Commonwealth of Australia, is a country in the Southern Hemisphere comprising the mainland of the Australian continent, the island of Tasmania, and numerous smaller islands in the Indian and Pacific Oceans. It is the world's sixth-largest country by total area...

 or perhaps neighboring Antarctica and did not reach Europe until less than 30 million years ago.

Unfortunately, it is unknown what the affiliations of the holotype
Holotype
A holotype is a single physical example of an organism, known to have been used when the species was formally described. It is either the single such physical example or one of several such, but explicitly designated as the holotype...

 material are. The fact that it was originally perceived to be something similar to a pratincole
Pratincole
The Pratincoles or Greywaders are a group of birds which together with the coursers and Egyptian Plover make up the family Glareolidae. They have short legs, very long pointed wings and long forked tails....

 or courser
Courser
The Coursers are a group of birds which together with the pratincoles make up the family Glareolidae. They have long legs, short wings and long pointed bills which curve downwards...

 suggests that the revised taxon
Taxon
|thumb|270px|[[African elephants]] form a widely-accepted taxon, the [[genus]] LoxodontaA taxon is a group of organisms, which a taxonomist adjudges to be a unit. Usually a taxon is given a name and a rank, although neither is a requirement...

 Precursor may belong into the Charadriiformes
Charadriiformes
Charadriiformes is a diverse order of small to medium-large birds. It includes about 350 species and has members in all parts of the world. Most Charadriiformes live near water and eat invertebrates or other small animals; however, some are pelagic , some occupy deserts and a few are found in thick...

. Certainly the Glareolidae
Glareolidae
Glareolidae is a family of birds in the wader suborder Charadri. It contains two distinct groups, the pratincoles and the coursers. The coursers include the atypical Egyptian Plover, Pluvianus aegyptius, which has sometimes been placed in its own family...

 are an ancient lineage, possibly originating more than 60 million years ago. If the "charadriimorph" remains are indeed Glareolidae, they should already be rather autapomorphic.
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