Mesoproterozoic
Encyclopedia
The Mesoproterozoic Era is a geologic
Geology
Geology is the science comprising the study of solid Earth, the rocks of which it is composed, and the processes by which it evolves. Geology gives insight into the history of the Earth, as it provides the primary evidence for plate tectonics, the evolutionary history of life, and past climates...

 era that occurred between 1600 Ma and 1000 Ma (million years ago). The Mesoproterozoic was the first period of Earth's history with a respectable geological record. Continents existed in the Paleoproterozoic
Paleoproterozoic
The Paleoproterozoic is the first of the three sub-divisions of the Proterozoic occurring between . This is when the continents first stabilized...

, but we know little about them. The continental masses of the Mesoproterozoic are more or less the same ones that are with us today.

The major events of this era are the formation of the Rodinia
Rodinia
In geology, Rodinia is the name of a supercontinent, a continent which contained most or all of Earth's landmass. According to plate tectonic reconstructions, Rodinia existed between 1.1 billion and 750 million years ago, in the Neoproterozoic era...

 supercontinent
Supercontinent
In geology, a supercontinent is a landmass comprising more than one continental core, or craton. The assembly of cratons and accreted terranes that form Eurasia qualifies as a supercontinent today.-History:...

, the breakup of the Columbia supercontinent
Columbia (supercontinent)
Columbia, also known as Nuna and Hudsonland, was one of Earth's oldest supercontinents. It was first proposed by J.J.W. Rogers and M. Santosh and is thought to have existed approximately 1.8 to 1.5 billion years ago in the Paleoproterozoic Era. Zhao et al...

, and the evolution of sexual reproduction
Sexual reproduction
Sexual reproduction is the creation of a new organism by combining the genetic material of two organisms. There are two main processes during sexual reproduction; they are: meiosis, involving the halving of the number of chromosomes; and fertilization, involving the fusion of two gametes and the...

.

This era is marked by the further development of continental plates and plate tectonics. At the end of this era, the continental plates that had developed were more or less the same we have today. This is the first era of which a good geological record still exists today.

The first large scale mountain building episode, the Grenville Orogeny
Grenville orogeny
The Grenville Orogeny was a long-lived Mesoproterozoic mountain-building event associated with the assembly of the supercontinent Rodinia. Its record is a prominent orogenic belt which spans a significant portion of the North American continent, from Labrador to Mexico, as well as to Scotland...

, for which extensive evidence still survives happened in this period.

This era was the high point of the Stromatolites before they declined in the Neoproterozoic
Neoproterozoic
The Neoproterozoic Era is the unit of geologic time from 1,000 to 542.0 ± 1.0 million years ago. The terminal Era of the formal Proterozoic Eon , it is further subdivided into the Tonian, Cryogenian, and Ediacaran Periods...

.

The era saw the development of sexual reproduction, which greatly increased the complexity of life to come. It was the start of development of communal living among organisms, the multicellular organisms.

It was an Era of apparently critical, but still poorly understood, changes in the chemistry of the sea, the sediments of the earth, and the composition of the air. Oxygen levels had risen to perhaps 1% of today's levels at the beginning of the era and continued rising throughout the Era.

Subdivisions

The subdivisions of the Mesoproterozoic are, obviously, arbitrary divisions based on time. They are not geo- or biostratigraphic units. The base of the Mesoproterozoic is defined chronometrically, in terms of years, rather than by the appearance or disappearance of some organism. This gives an illusory sense of certainty. Radiometric dating
Radiometric dating
Radiometric dating is a technique used to date materials such as rocks, usually based on a comparison between the observed abundance of a naturally occurring radioactive isotope and its decay products, using known decay rates...

 is a good tool, and gets better each decade. This creates certain problems. As a practical matter, radiometric dates have an error margin of 1-2%. That sounds good, but it means that two sites, both measured to be at the exact base of the Ectasian
Ectasian
The Ectasian is the second geologic period in the Mesoproterozoic Era and lasted from 1400 Mya ago to 1200 Mya...

, might differ in age by over 50 My. Since the Ectasian is only 200 My long, that's a serious matter. And this accounts only for random error. Systematic errors can be caused by extraterrestrial events, by geochemical or biochemical sorting of isotopes, and human error. Thus far, biostratigraphy
Biostratigraphy
Biostratigraphy is the branch of stratigraphy which focuses on correlating and assigning relative ages of rock strata by using the fossil assemblages contained within them. Usually the aim is correlation, demonstrating that a particular horizon in one geological section represents the same period...

 has usually proved considerably more exact. In addition, a thoughtful choice of biological marker can be used as a signal to expect a whole host of ecological changes. The difference between a Changhsingian
Changhsingian
In the geologic timescale, the Changhsingian or Changxingian is the latest age or uppermost stage of the Permian. It is also the upper or latest of two subdivisions of the Lopingian epoch or series. The Changhsingian lasted from 253.8 ± 0.7 to 251.0 ± 0.7 million years ago...

 and an Induan
Induan
The Induan is, in the geologic timescale, the first age of the Early Triassic epoch or the lowest stage of the Lower Triassic series. It spans the time between 251 ± 0.4 Ma and 249.7 ± 0.7 Ma...

 deposit isn't just a matter of a few years. The world changed hugely at the end of the Permian.

By contrast, the transition from Calymmian
Calymmian
The Calymmian is the first geologic period in the Mesoproterozoic Era and lasted from 1600 Mya to 1400 Mya...

to Ectasian has no meaning beyond calendar time. The usual reason given for the use of a chronometric system is that there is insufficient biological activity or geochemical change to find useful markers. That is a position which is now a little uncertain and is going to become increasingly tenuous over the next few years. For example, there are a number of good potential markers in the rise and decline of "Christmas tree" stromatolites, in the coming and going of banded iron formations, the appearance of stable carbon-13 isotope (13C) excursions, and so on. These have real meaning for the geologist and paleontologist.

For that matter, they are not completely without biological markers. There has been considerable progress in studying and identifying fossil bacteria and Eukarya. The cyanobacterium Archaeoellipsoides is one relatively common form, apparently known from several species. It is probably related to the extant Anabaena and indicates the presence of significant free oxygen. Oxygen levels also had significant effects on ocean chemistry: increasing continental weathering rates and providing sulfates and nitrates as nutrients. It would be remarkable if this didn't result, in turn in new populations of both bacterial and eukaryotic organisms. Since the presence of these cells would be tied directly to important geochemical events, they would make ideal organisms for biostratigraphy.

Clearly, clocks cannot be set by hypothetical organisms depending on, as yet, poorly understood chemistry. Not enough is known about these matters yet.

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