Lonchodytes is a Late Cretaceous

Late Cretaceous

The Late Cretaceous is the younger of two epochs into which the Cretaceous period is divided in the geologic timescale. Rock strata from this epoch form the Upper Cretaceous series...

 genus

Genus

In biology, a genus is a low-level taxonomic rank used in the biological classification of living and fossil organisms, which is an example of definition by genus and differentia...

 of aquatic

Aquatic animal

An aquatic animal is an animal, either vertebrate or invertebrate, which lives in water for most or all of its life. It may breathe air or extract its oxygen from that dissolved in water through specialised organs called gills, or directly through its skin. Natural environments and the animals that...

 bird

Bird

Birds are feathered, winged, bipedal, endothermic , egg-laying, vertebrate animals. Around 10,000 living species and 188 families makes them the most speciose class of tetrapod vertebrates. They inhabit ecosystems across the globe, from the Arctic to the Antarctic. Extant birds range in size from...

, which lived along the shores of the Western Interior Seaway

Western Interior Seaway

The Western Interior Seaway, also called the Cretaceous Seaway, the Niobraran Sea, and the North American Inland Sea, was a huge inland sea that split the continent of North America into two halves, Laramidia and Appalachia, during most of the mid- and late-Cretaceous Period...

. It lived probably during the Maastrichtian

Maastrichtian

The Maastrichtian is, in the ICS' geologic timescale, the latest age or upper stage of the Late Cretaceous epoch or Upper Cretaceous series, the Cretaceous period or system, and of the Mesozoic era or erathem. It spanned from 70.6 ± 0.6 Ma to 65.5 ± 0.3 Ma...

, 70 million years ago (mya), and was found in Lance Creek Formation rocks in Wyoming

Wyoming

Wyoming is a state in the mountain region of the Western United States. The western two thirds of the state is covered mostly with the mountain ranges and rangelands in the foothills of the Eastern Rocky Mountains, while the eastern third of the state is high elevation prairie known as the High...

 (USA) though it seems still somewhat unclear if it did fossil

Fossil

Fossils are the preserved remains or traces of animals , plants, and other organisms from the remote past...

ize there or was reworked from later (Danian

Danian

The Danian is the oldest age or lowermost stage of the Paleocene epoch or series, the Paleogene period or system and the Cenozoic era or erathem. The beginning of the Danian age is at the Cretaceous-Tertiary extinction event...

: Early Paleocene

Paleocene

The Paleocene or Palaeocene, the "early recent", is a geologic epoch that lasted from about . It is the first epoch of the Palaeogene Period in the modern Cenozoic Era...

, less than 65 mya) deposits.

Lonchodytes estesi, the type species

Type species

In biological nomenclature, a type species is both a concept and a practical system which is used in the classification and nomenclature of animals and plants. The value of a "type species" lies in the fact that it makes clear what is meant by a particular genus name. A type species is the species...

, appears to be closely related to the ancestor of some modern birds. It is most often allied - albeit tentatively - with the Gaviiformes

Gaviiformes

Gaviiformes is an order of aquatic birds containing the loons or divers and their closest extinct relatives. Modern gaviiformes are found in many parts of North America and northern Eurasia , though prehistoric species were more widespread.-Classification and evolution:There are five living...

 (loons/divers), Procellariiformes

Procellariiformes

Procellariiformes is an order of seabirds that comprises four families: the albatrosses, petrels and shearwaters, storm petrels, and diving petrels...

 (tubenoses) or Pelecaniformes

Pelecaniformes

The Pelecaniformes is a order of medium-sized and large waterbirds found worldwide. As traditionally—but erroneously—defined, they encompass all birds that have feet with all four toes webbed. Hence, they were formerly also known by such names as totipalmates or steganopodes...

. It was found to be closer to Procellariiformes than to the other order

Order (biology)

In scientific classification used in biology, the order is# a taxonomic rank used in the classification of organisms. Other well-known ranks are life, domain, kingdom, phylum, class, family, genus, and species, with order fitting in between class and family...

s by a cladistic analysis

These are today generally believed to be closely related if not actually a clade

Clade

A clade is a group consisting of a species and all its descendants. In the terms of biological systematics, a clade is a single "branch" on the "tree of life". The idea that such a "natural group" of organisms should be grouped together and given a taxonomic name is central to biological...

, perhaps together with other "higher waterbirds" such as penguin

Penguin

Penguins are a group of aquatic, flightless birds living almost exclusively in the southern hemisphere, especially in Antarctica. Highly adapted for life in the water, penguins have countershaded dark and white plumage, and their wings have become flippers...

s. L. estesi might be basal to some lineages among these three, and/or other "higher waterbirds", a hypothesis by and large compatible with current phylogenetic analyses based on morphological

Morphology (biology)

In biology, morphology is a branch of bioscience dealing with the study of the form and structure of organisms and their specific structural features....

 and DNA sequence

DNA sequence

The sequence or primary structure of a nucleic acid is the composition of atoms that make up the nucleic acid and the chemical bonds that bond those atoms. Because nucleic acids, such as DNA and RNA, are unbranched polymers, this specification is equivalent to specifying the sequence of...

 data, as well as the Mesozoic

Mesozoic

The Mesozoic era is an interval of geological time from about 250 million years ago to about 65 million years ago. It is often referred to as the age of reptiles because reptiles, namely dinosaurs, were the dominant terrestrial and marine vertebrates of the time...

 record of Neornithes

"Lonchodytes" pterygius on the other hand quite likely does not belong into this genus. It appears very distinct from in a cladistic phylogeny and seems to be a member of the Charadriiformes

Charadriiformes

Charadriiformes is a diverse order of small to medium-large birds. It includes about 350 species and has members in all parts of the world. Most Charadriiformes live near water and eat invertebrates or other small animals; however, some are pelagic , some occupy deserts and a few are found in thick...

. Given the paucity of remains however, it cannot be excluded with certainty that it represents an ancestral "higher waterbird" lineage that was morphologically

Morphology (biology)

In biology, morphology is a branch of bioscience dealing with the study of the form and structure of organisms and their specific structural features....

 convergent to waders.