Lombricine is a phosphagen
The phosphagens are energy storage compounds, also known as high-energy phosphate compounds, are chiefly found in muscular tissue in animals. They allow a high-energy phosphate pool to be maintained in a concentration range, which, if it all were ATP, would create problems due to the ATP consuming...

 that is unique to earthworms. Structurally, it is a phosphodiester of the guanidine
Guanidine is a crystalline compound of strong alkalinity formed by the oxidation of guanine. It is used in the manufacture of plastics and explosives. It is found in urine as a normal product of protein metabolism. The molecule was first synthesized in 1861 by the oxidative degradation of an...

, 2-guanidinoethanol (IUPAC name: 2-carbamimidamidoethanol), and D-serine (not the usual L-serine), which is then further phosphorylated by lombricine kinase
Lombricine kinase
In enzymology, a lombricine kinase is an enzyme that catalyzes the chemical reactionThus, the two substrates of this enzyme are ATP and lombricine, whereas its two products are ADP and N-phospholombricine....

 to phospholombricine.
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