Hadrianus (genus)
Encyclopedia
Hadrianus is an extinct genus of tortoise
Tortoise
Tortoises are a family of land-dwelling reptiles of the order of turtles . Like their marine cousins, the sea turtles, tortoises are shielded from predators by a shell. The top part of the shell is the carapace, the underside is the plastron, and the two are connected by the bridge. The tortoise...

 belonging to the Testudinidae
Family (biology)
In biological classification, family is* a taxonomic rank. Other well-known ranks are life, domain, kingdom, phylum, class, order, genus, and species, with family fitting between order and genus. As for the other well-known ranks, there is the option of an immediately lower rank, indicated by the...

 found in the United States
United States
The United States of America is a federal constitutional republic comprising fifty states and a federal district...

 and Spain
Spain
Spain , officially the Kingdom of Spain languages]] under the European Charter for Regional or Minority Languages. In each of these, Spain's official name is as follows:;;;;;;), is a country and member state of the European Union located in southwestern Europe on the Iberian Peninsula...

 and believed to be the oldest true tortoise known. The genus is thought to be closely related to the genus Manouria
Manouria
Manouria is a genus of tortoise in the Testudinidae family.It contains the following species:* Impressed Tortoise * Manouria emys* Manouria sondaari - an extinct giant land tortoise from Luzon Island, Philippines....

. The genus may have evolved in the subtropics of Asia and subsequently migrated to North America and Europe.
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