The Four-spotted Butterflyfish or fourspot butterflyfish (Chaetodon quadrimaculatus) is a species

Species

In biology, a species is one of the basic units of biological classification and a taxonomic rank. A species is often defined as a group of organisms capable of interbreeding and producing fertile offspring. While in many cases this definition is adequate, more precise or differing measures are...

 of butterflyfish

Butterflyfish

The butterflyfish are a group of conspicuous tropical marine fish of the family Chaetodontidae; the bannerfish and coralfish are also included in this group. Found mostly on the reefs of the Atlantic, Indian and Pacific Oceans, there are approximately 120 species in 10 genera...

 (family

Family (biology)

In biological classification, family is* a taxonomic rank. Other well-known ranks are life, domain, kingdom, phylum, class, order, genus, and species, with family fitting between order and genus. As for the other well-known ranks, there is the option of an immediately lower rank, indicated by the...

 Chaetodontidae) found in the Pacific Ocean

Pacific Ocean

The Pacific Ocean is the largest of the Earth's oceanic divisions. It extends from the Arctic in the north to the Southern Ocean in the south, bounded by Asia and Australia in the west, and the Americas in the east.At 165.2 million square kilometres in area, this largest division of the World...

 from the Ryukyus, Ogasawara

Ogasawara Islands

The Bonin Islands, known in Japan as the are an archipelago of over 30 subtropical and tropical islands, some directly south of Tokyo, Japan. Administratively, they are part of Ogasawara Municipality of Ogasawara Subprefecture, Tokyo...

 (Bonin) Islands and Taiwan

Taiwan

Taiwan , also known, especially in the past, as Formosa , is the largest island of the same-named island group of East Asia in the western Pacific Ocean and located off the southeastern coast of mainland China. The island forms over 99% of the current territory of the Republic of China following...

 to the Hawaiian

Hawaiian Islands

The Hawaiian Islands are an archipelago of eight major islands, several atolls, numerous smaller islets, and undersea seamounts in the North Pacific Ocean, extending some 1,500 miles from the island of Hawaii in the south to northernmost Kure Atoll...

, Marquesan

Marquesas Islands

The Marquesas Islands enana and Te Fenua `Enata , both meaning "The Land of Men") are a group of volcanic islands in French Polynesia, an overseas collectivity of France in the southern Pacific Ocean. The Marquesas are located at 9° 00S, 139° 30W...

, and Pitcairn islands, south to the Samoa

Samoa

Samoa , officially the Independent State of Samoa, formerly known as Western Samoa is a country encompassing the western part of the Samoan Islands in the South Pacific Ocean. It became independent from New Zealand in 1962. The two main islands of Samoa are Upolu and one of the biggest islands in...

n and Austral Islands

Austral Islands

The Austral Islands are the southernmost group of islands in French Polynesia, an overseas collectivity of France in the South Pacific. Geographically, they consist of two separate archipelagos, namely in the northwest the Tubuai Islands consisting of the Îles Maria, Rimatara, Rurutu, Tubuai...

 and the Marianas and Marshall Islands

Marshall Islands

The Republic of the Marshall Islands , , is a Micronesian nation of atolls and islands in the middle of the Pacific Ocean, just west of the International Date Line and just north of the Equator. As of July 2011 the population was 67,182...

 in Micronesia

Micronesia

Micronesia is a subregion of Oceania, comprising thousands of small islands in the western Pacific Ocean. It is distinct from Melanesia to the south, and Polynesia to the east. The Philippines lie to the west, and Indonesia to the southwest....

.

It is a quite distinct species, but most closely related to the Speckled Butterflyfish

Speckled butterflyfish

Chaetodon citrinellus is a species of butterflyfish . It is commonly known as the Speckled Butterflyfish or Citron Butterflyfish...

 (C. citrinellus). Together they are basal in the subgenus

Subgenus

In biology, a subgenus is a taxonomic rank directly below genus.In zoology, a subgeneric name can be used independently or included in a species name, in parentheses, placed between the generic name and the specific epithet: e.g. the Tiger Cowry of the Indo-Pacific, Cypraea tigris Linnaeus, which...

 Exornator, and might be intermediate between the core group of this subgenus and the species of the Rhombochaetodon (or Roaops) lineage. If that is correct, the latter would require to be merged into Exornator. If the genus Chaetodon

Chaetodon

Chaetodon is a tropical fish genus in the family Chaetodontidae. Like their relatives, they are known as "butterflyfish". This genus is by far the largest among the Chaetodontidae, with about 90 living species included here, though most might warrant recognition as distinct genera.Several subgenera...

is split up, Exornator might become a subgenus of Lepidochaetodon.