Folke K Skoog
Encyclopedia
Folke Karl Skoog was a Swedish
Sweden
Sweden , officially the Kingdom of Sweden , is a Nordic country on the Scandinavian Peninsula in Northern Europe. Sweden borders with Norway and Finland and is connected to Denmark by a bridge-tunnel across the Öresund....

-born American plant physiologist
Plant physiology
Plant physiology is a subdiscipline of botany concerned with the functioning, or physiology, of plants. Closely related fields include plant morphology , plant ecology , phytochemistry , cell biology, and molecular biology.Fundamental processes such as photosynthesis, respiration, plant nutrition,...

 who was a pioneer in the field of plant growth regulators, particularly cytokinins. Skoog was a recipient of the National Medal of Science
National Medal of Science
The National Medal of Science is an honor bestowed by the President of the United States to individuals in science and engineering who have made important contributions to the advancement of knowledge in the fields of behavioral and social sciences, biology, chemistry, engineering, mathematics and...

.

Born in Halland, Sweden
Sweden
Sweden , officially the Kingdom of Sweden , is a Nordic country on the Scandinavian Peninsula in Northern Europe. Sweden borders with Norway and Finland and is connected to Denmark by a bridge-tunnel across the Öresund....

, Skoog emigrated to the United States during a trip to California
California
California is a state located on the West Coast of the United States. It is by far the most populous U.S. state, and the third-largest by land area...

 in 1925, and was naturalized as a citizen almost a decade later. He competed, and finished sixth in heat 2, in the 1500 meter race during the 1932 Summer Olympics
1932 Summer Olympics
The 1932 Summer Olympics, officially known as the Games of the X Olympiad, was a major world wide multi-athletic event which was celebrated in 1932 in Los Angeles, California, United States. No other cities made a bid to host these Olympics. Held during the worldwide Great Depression, many nations...

. In 1936, he received his PhD in biology from Caltech for his work done with auxin
Auxin
Auxins are a class of plant hormones with some morphogen-like characteristics. Auxins have a cardinal role in coordination of many growth and behavioral processes in the plant's life cycle and are essential for plant body development. Auxins and their role in plant growth were first described by...

, a plant hormone.

Skoog's professional career advanced significantly with his arrival at the University of Wisconsin–Madison
University of Wisconsin–Madison
The University of Wisconsin–Madison is a public research university located in Madison, Wisconsin, United States. Founded in 1848, UW–Madison is the flagship campus of the University of Wisconsin System. It became a land-grant institution in 1866...

 in 1947. Carlos Miller discovered kinetin
Kinetin
Kinetin is a kind of cytokinin, a class of plant hormone that promotes cell division. Kinetin was originally isolated by Miller and Skoog et al. as a compound from autoclaved herring sperm DNA that had cell division-promoting activity. It was given the name kinetin because of its ability to induce...

 in 1954, and benzyladenine and related compounds were later synthesized in Skoog's lab.

In 1962, Skoog and Toshio Murashige
Toshio Murashige
Toshio Murashige is a professor emeritus of University of California Riverside in plant biology.He is most widely known for his efforts in creating the plant tissue culture medium known as Murashige and Skoog medium.- External links :*...

 published what is probably the best-known paper in plant tissue culture; in a fruitless attempt to discover a yet-unknown plant growth regulator in tobacco
Tobacco
Tobacco is an agricultural product processed from the leaves of plants in the genus Nicotiana. It can be consumed, used as a pesticide and, in the form of nicotine tartrate, used in some medicines...

 juice for his doctoral thesis, Murashige and Skoog instead developed a greatly improved salt base for the sterile culture of tobacco. Now referred to as Murashige and Skoog medium
Murashige and Skoog medium
Murashige and Skoog medium or is a plant growth medium used in the laboratories for cultivation of plant cell culture. MSO was invented by plant scientists Toshio Murashige and Folke K. Skoog in 1962 during Murashige's search for a new plant growth regulator...

, the final paper (Murashige, T. and Skoog, F. (1962) A revised medium for rapid growth and bioassays with tobacco tissue cultures. Physiol Plant 18: 100-127) is considered one of the most often-cited papers in biology. Now 50 years after the work, M&S salt base remains an essential component in plant tissue culture
Plant tissue culture
Plant tissue culture is a collection of techniques used to maintain or grow plant cells, tissues or organs under sterile conditions on a nutrient culture medium of known composition. Plant tissue culture is widely used to produce clones of a plant in a method known as micropropagation...

.

In 1970, Skoog was elected a foreign member of the Royal Swedish Academy of Sciences
Royal Swedish Academy of Sciences
The Royal Swedish Academy of Sciences or Kungliga Vetenskapsakademien is one of the Royal Academies of Sweden. The Academy is an independent, non-governmental scientific organization which acts to promote the sciences, primarily the natural sciences and mathematics.The Academy was founded on 2...

.
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