Drilliidae
Encyclopedia
Drilliidae is a taxonomic family
of small sea snail
s, marine
gastropod mollusks in the superfamily Conoidea.
This family has no subfamilies.
is truncated. The aperture
is U-shaped with a parietal callus pad. The sculpture of the shell shows prominent axial ribs with a polished surface. Most species have a dorsal varix
(transverse elevation), except in the genera Cymatosyrinx
, Elaeocyma
and Splendrillia
. The protoconch
can be smooth or very carinate. The ovate operculum
has a terminal nucleus.
The radula
of the species in this family have characteristically five teeth in each row (formula : 1 + 1 + 1 + 1 + 1) with a vestigial central tooth, comb-like lateral teeth and a pair of flat-pointed, slender marginal teeth.
The foregut
structure of the anterior alimentary system has the least derived foregut anatomy of all the families in the superfamily Conoidea
.
with comb-like lateral teeth. Finally, in 1993, Taylor et al. proposed the promotion of Drillinae fom subfamily to the family level Drilliidae.
Genera brought into synonymy:
Genera moved to other families:
Family (biology)
In biological classification, family is* a taxonomic rank. Other well-known ranks are life, domain, kingdom, phylum, class, order, genus, and species, with family fitting between order and genus. As for the other well-known ranks, there is the option of an immediately lower rank, indicated by the...
of small sea snail
Sea snail
Sea snail is a common name for those snails that normally live in saltwater, marine gastropod molluscs....
s, marine
Marine (ocean)
Marine is an umbrella term. As an adjective it is usually applicable to things relating to the sea or ocean, such as marine biology, marine ecology and marine geology...
gastropod mollusks in the superfamily Conoidea.
This family has no subfamilies.
Description
The shell is claviform with a tall spire , or squatly conical to biconical. In many species, the siphonal canalSiphonal canal
Some sea marine gastropods have a soft tubular anterior extension of the mantle called a siphon through which water is drawn into the mantle cavity and over the gill and which serves as a chemoreceptor to locate food. In many carnivorous snails, where the siphon is particularly long, the structure...
is truncated. The aperture
Aperture (mollusc)
The aperture is an opening in certain kinds of mollusc shells: it is the main opening of the shell, where part of the body of the animal emerges for locomotion, feeding, etc....
is U-shaped with a parietal callus pad. The sculpture of the shell shows prominent axial ribs with a polished surface. Most species have a dorsal varix
Varix (mollusc)
A varix is an anatomical feature of the shell of certain sea snails, marine gastropod molluscs.It is a thickened axial ridge in the shell of some families of gastropods. A varix is located at intervals around the whorl, and is formed by considerable thickening of the outer lip at a resting stage...
(transverse elevation), except in the genera Cymatosyrinx
Cymatosyrinx
Cymatosyrinx is a genus of sea snails, marine gastropod mollusks in the family Drilliidae. -Species:Species within the genus Cymatosyrinx include:* Cymatosyrinx arbela * Cymatosyrinx bartschi Haas, 1941...
, Elaeocyma
Elaeocyma
Elaeocyma is a genus of sea snails, marine gastropod mollusks in the family Drilliidae. -Species:Species within the genus Elaeocyma include:* Elaeocyma amplinucis McLean & Poorman, 1971* Elaeocyma arenensis...
and Splendrillia
Splendrillia
Splendrillia is a genus of sea snails, marine gastropod mollusks in the family Drilliidae. -Species:Species within the genus Splendrillia include:* Splendrillia academica McLean & Poorman, 1971* Splendrillia alabastrum Kilburn, 1988...
. The protoconch
Protoconch
A protoconch is an embryonic or larval shell of some classes of molluscs, e.g., the initial chamber of an ammonite or the larval shell of a gastropod...
can be smooth or very carinate. The ovate operculum
Operculum (gastropod)
The operculum, meaning little lid, is a corneous or calcareous anatomical structure which exists in many groups of sea snails and freshwater snails, and also in a few groups of land snails...
has a terminal nucleus.
The radula
Radula
The radula is an anatomical structure that is used by molluscs for feeding, sometimes compared rather inaccurately to a tongue. It is a minutely toothed, chitinous ribbon, which is typically used for scraping or cutting food before the food enters the esophagus...
of the species in this family have characteristically five teeth in each row (formula : 1 + 1 + 1 + 1 + 1) with a vestigial central tooth, comb-like lateral teeth and a pair of flat-pointed, slender marginal teeth.
The foregut
Foregut
The foregut is the anterior part of the alimentary canal, from the mouth to the duodenum at the entrance of the bile duct. At this point it is continuous with the midgut...
structure of the anterior alimentary system has the least derived foregut anatomy of all the families in the superfamily Conoidea
Conoidea
Conoidea is a superfamily of predatory sea snails, marine gastropod mollusks within the suborder Hypsogastropoda. This superfamily is a very large group of marine mollusks, estimated at about 340 recent valid genera and subgenera, and considered by one authority to contain 4,000 named living species...
.
Taxonomic history
The genera in this family were originally separated from the subfamily Clavitulinae and classified by H.& A. Adams (1858) under subfamily Turritinae, because their operculum has a terminal instead of a central nucleus. In 1942 the species with a U-shaped sinus and a parietal callus pad were brought by Powell in the new subfamily Clavinae. In 1966 Morrison proposed the subfamily Drillinae ( = Clavinae) for the species with a stenoglossan radulaRadula
The radula is an anatomical structure that is used by molluscs for feeding, sometimes compared rather inaccurately to a tongue. It is a minutely toothed, chitinous ribbon, which is typically used for scraping or cutting food before the food enters the esophagus...
with comb-like lateral teeth. Finally, in 1993, Taylor et al. proposed the promotion of Drillinae fom subfamily to the family level Drilliidae.
Taxonomy
Genera in the family Drilliidae include:- ? AcinodrilliaAcinodrilliaAcinodrillia is a genus of sea snails, marine gastropod mollusks in the family Drilliidae. -Species:Species within the genus Acinodrillia include:* Acinodrillia amazimba Kilburn, 1988* Acinodrillia paula...
Kilburn, 1988 - AgladrilliaAgladrilliaAgladrillia is a genus of sea snails, marine gastropod mollusks in the family Drilliidae. -Species:Species within the genus Agladrillia include:* Agladrillia anadelgado Rolán et al., 2007* Agladrillia badia McLean & Poorman, 1971...
Woodring, 1928 - BellaspiraBellaspiraBellaspira is a genus of sea snails, marine gastropod mollusks in the family Drilliidae. -Species:Species within the genus Bellaspira include:* Bellaspira acclivicosta McLean & L. Poorman, 1970* Bellaspira brunnescens...
Conrad, 1868 - CalliclavaCalliclavaCalliclava is a genus of sea snails, marine gastropod mollusks in the family Drilliidae. -Species:Species within the genus Calliclava include:* Calliclava aegina * Calliclava albolaqueata...
McLean, 1971 - CerodrilliaCerodrilliaCerodrillia is a genus of sea snails, marine gastropod mollusks in the family Drilliidae. -Species:Species within the genus Cerodrillia include:* Cerodrillia asymmetrica McLean & Poorman, 1971* Cerodrillia bahamensis...
Bartsch & Rehder, 1939 - ClathrodrilliaClathrodrilliaClathrodrillia is a genus of sea snails, marine gastropod mollusks in the family Drilliidae. -Species:Species within the genus Clathrodrillia include:* Clathrodrillia albicoma...
Dall, 1918 - ClavusClavus (gastropod)Clavus is a genus of small sea snails, marine gastropod mollusks in the family Drilliidae. -Description:Apart from the general characteristics of the genera in the family Drilliidae, the species in the genus Clavus have a shell with peripheral tubercles, spines or wing-like processes. The...
Montfort, 1810 - ConopleuraConopleuraConopleura is a genus of sea snails, marine gastropod mollusks in the family Drilliidae. -Species:Species within the genus Conopleura include:* Conopleura aliena Smriglio, Mariottini & Calascibetta, 1999* Conopleura striata Hinds, 1844...
Hinds, 1844 - Crassopleura Monterosato, 1884
- ? Cruziturricula Marks, 1951
- CymatosyrinxCymatosyrinxCymatosyrinx is a genus of sea snails, marine gastropod mollusks in the family Drilliidae. -Species:Species within the genus Cymatosyrinx include:* Cymatosyrinx arbela * Cymatosyrinx bartschi Haas, 1941...
Dall, 1889 - DouglassiaDouglassiaDouglassia is a genus of sea snails, marine gastropod mollusks in the family Drilliidae.-Species:Species within the genus Douglassia include:...
Bartsch, 1934 - DrilliaDrilliaDrillia is a genus of small sea snails, marine gastropod mollusks in the family Drilliidae. -General characteristics:The shell is turriculated, with an elevated spire, longitudinal ribs, and usually revolving striae. The spiral sculpture of the shell is rather smooth or shows a wide spacing of...
Gray, 1838 - ElaeocymaElaeocymaElaeocyma is a genus of sea snails, marine gastropod mollusks in the family Drilliidae. -Species:Species within the genus Elaeocyma include:* Elaeocyma amplinucis McLean & Poorman, 1971* Elaeocyma arenensis...
Dall, 1918 - EumetadrilliaEumetadrilliaEumetadrillia is a genus of sea snails, marine gastropod mollusks in the family Drilliidae.-Species:Species within the genus Eumetadrillia include:* † Eumetadrillia acidna Woodring, 1970* † Eumetadrillia dodona Gardner, 1937...
Woodring, 1928 - FenimoreaFenimoreaFenimorea is a genus of sea snails, marine gastropod mollusks in the family Drilliidae. -Species:Species within the genus Fenimorea include:* Fenimorea culexensis Nowell-Usticke, 1969* Fenimorea fucata...
Bartsch, 1934 - ? FusiturriculaFusiturriculaFusiturricula is a genus of sea snails, marine gastropod mollusks in the family Drilliidae. -Taxonomy:The genus Fusiturricula forms an unsupported group that is sister clade to Drilliidae in the cladogram of the molecular phylogeny of the Conoidea....
Woodring, 1928 - GlobidrilliaGlobidrilliaGlobidrillia is a genus of sea snails, marine gastropod mollusks in the family Drilliidae. -Species:Species within the genus Globidrillia include:* Globidrillia ferminiana * Globidrillia hemphillii...
Woodring, 1928 - Hauturua Powell, 1942
- ImaclavaImaclavaImaclava is a genus of sea snails, marine gastropod mollusks in the family Drilliidae. -Species:Species within the genus Imaclava include:* Imaclava asaedai * Imaclava hotei...
Bartsch, 1944 - IredaleaIredaleaIredalea is a small genus of sea snails, marine gastropod mollusks in the family Drilliidae.This genus was named in 1915 by Walter Oliver after Tom Iredale , an English-born naturalist who lived mainly in Australia.-Species:...
Oliver, 1915 - KylixKylix (gastropod)Kylix is a genus of sea snails, marine gastropod mollusks in the family Drilliidae. -Species:Species within the genus Kylix include:* Kylix alcyone * Kylix contracta McLean & Poorman, 1971* Kylix halocydne...
Dall, 1919 - LeptadrilliaLeptadrilliaLeptadrillia is a genus of sea snails, marine gastropod mollusks in the family Drilliidae. -Species:Species within the genus Leptadrillia include:* Leptadrillia cinereopellis Kuroda, Habe & Oyama, 1971* Leptadrillia cookei Leptadrillia is a genus of sea snails, marine gastropod mollusks in the...
Woodring, 1928 - Lissodrillia Bartsch & Rehder, 1943 (considered as a subspecies of CerodrilliaCerodrilliaCerodrillia is a genus of sea snails, marine gastropod mollusks in the family Drilliidae. -Species:Species within the genus Cerodrillia include:* Cerodrillia asymmetrica McLean & Poorman, 1971* Cerodrillia bahamensis...
) - NeodrilliaNeodrilliaNeodrillia is a genus of sea snails, marine gastropod mollusks in the family Drilliidae. -Species:Species within the genus Neodrillia include:* Neodrillia blacki Petuch, 2003* Neodrillia cydia Bartsch, 1943...
Bartsch, 1943 - OrrmaesiaOrrmaesiaOrrmaesia is a genus of sea snails, marine gastropod mollusks in the family Drilliidae. -Species:Species within the genus Orrmaesia include:* Orrmaesia ancilla * Orrmaesia dorsicosta Kilburn, 1988...
Kilburn, 1988 - ? ParacuneusParacuneusParacuneus is a genus of sea snails, marine gastropod mollusks in the family Drilliidae.This genus is placed tentatively in the family Drilliidae. -Species:Species within the genus Paracuneus include:...
Laseron, 1954 - PlagiostrophaPlagiostrophaPlagiostropha is a genus of sea snails, marine gastropod mollusks in the family Drilliidae. -Species:Species within the genus Plagiostropha include:* Plagiostropha bicolor Chino & Stahlschmidt, 2010...
Melvill, 1927 - SpirotropisSpirotropis (gastropod)Spirotropis is a genus of sea snails, marine gastropod mollusks in the family Drilliidae. -Species:Species within the genus Spirotropis include:* Spirotropis agamedea * Spirotropis aganactica...
Sars, 1878 - SplendrilliaSplendrilliaSplendrillia is a genus of sea snails, marine gastropod mollusks in the family Drilliidae. -Species:Species within the genus Splendrillia include:* Splendrillia academica McLean & Poorman, 1971* Splendrillia alabastrum Kilburn, 1988...
Hedley, 1922 - ? StenodrilliaStenodrilliaStenodrillia is a genus of sea snails, marine gastropod mollusks in the family Drilliidae. -Species:Species within the genus Stenodrillia include:* Stenodrillia horrenda -References:...
Korobkov, 1955 - SyntomodrilliaSyntomodrilliaSyntomodrillia is a genus of sea snails, marine gastropod mollusks in the family Drilliidae. -Species:Species within the genus Syntomodrillia include:* Syntomodrillia vitrea McLean & Poorman, 1971...
Hedley, 1922 - TylotiellaTylotiellaTylotiella is a genus of sea snails, marine gastropod mollusks in the family Drilliidae. -Species:Species within the genus Tylotiella include:* Tylotiella androyensis Bozzetti, 2007* Tylotiella basipunctata Kilburn, 1988...
Habe, 1958
Genera brought into synonymy:
- BrephodrilliaBrephodrilliaBrephodrillia is a genus of sea snails, marine gastropod mollusks in the family Drilliidae.-Taxonomy:H. Pilsbry and H. Lowe put in Brephodrillia the species of Iredalea with a dorsal varix. with Brephodrillia perfectus Pilsbry and Lowe, 1932 as type species...
Pilsbry & Lowe, 1932: synonym of IredaleaIredaleaIredalea is a small genus of sea snails, marine gastropod mollusks in the family Drilliidae.This genus was named in 1915 by Walter Oliver after Tom Iredale , an English-born naturalist who lived mainly in Australia.-Species:... - Clavicantha Swainson, 1840 : synonym of ClavusClavusClavus may refer to:*Clavus : a genus of gastropods in the family Drilliidae*the plant disease ergot;*the Roman laticlave;*a shooting pain in the forehead, associated with hysteria, also called clavus hystericus;...
Montfort, 1810 - Eldridgea Bartsch, 1934 : synonym of ClavusClavusClavus may refer to:*Clavus : a genus of gastropods in the family Drilliidae*the plant disease ergot;*the Roman laticlave;*a shooting pain in the forehead, associated with hysteria, also called clavus hystericus;...
Montfort, 1810 - Fusisyrinx Bartsch, 1934 : synonym of FusiturriculaFusiturriculaFusiturricula is a genus of sea snails, marine gastropod mollusks in the family Drilliidae. -Taxonomy:The genus Fusiturricula forms an unsupported group that is sister clade to Drilliidae in the cladogram of the molecular phylogeny of the Conoidea....
Woodring, 1928 - Tylotia Melvill, 1917 : synonym of ClavusClavusClavus may refer to:*Clavus : a genus of gastropods in the family Drilliidae*the plant disease ergot;*the Roman laticlave;*a shooting pain in the forehead, associated with hysteria, also called clavus hystericus;...
Montfort, 1810
Genera moved to other families:
- AustroclavusAustroclavusAustroclavus is a genus of sea snails, marine gastropod mollusks, unassigned to a family within the superfamily Conoidea. -Species:Species within the genus Austroclavus include:* Austroclavus undata...
(unassigned to a family within Conoidea) - BrachytomaBrachytomaBrachytoma is a genus of sea snails, marine gastropod mollusks in the family Pseudomelatomidae. -Species:Species within the genus Brachytoma include:* Brachytoma rioensis * Brachytoma rufolineata...
moved to the family PseudomelatomidaePseudomelatomidaePseudomelatomidae is a family of predatory sea snails, marine gastropods included in the superfamily Conoidea and part of the Neogastropoda... - SediliopsisSediliopsisSediliopsis is a genus of sea snails, marine gastropod mollusks in the family Drilliidae.-Species:Species within the genus Sediliopsis include:* Sediliopsis riosi Tippett, 1995...
Petuch, 1988 : moved to the family PseudomelatomidaePseudomelatomidaePseudomelatomidae is a family of predatory sea snails, marine gastropods included in the superfamily Conoidea and part of the Neogastropoda...