Domain (biology)
Encyclopedia
In biological
taxonomy
, a domain (also superregnum, superkingdom, empire, or regio) is the highest taxonomic rank
of organism
s, higher than a kingdom
. According to the three-domain system
of Carl Woese
, introduced in 1990, the Tree of Life
consists of three domains: Archaea
, Bacteria
and Eukarya. The arrangement of taxa
reflects the fundamental differences in the genome
s.
Alternative classifications of life
so far proposed include:
None of the three systems currently include non-cellular life
.
Biology
Biology is a natural science concerned with the study of life and living organisms, including their structure, function, growth, origin, evolution, distribution, and taxonomy. Biology is a vast subject containing many subdivisions, topics, and disciplines...
taxonomy
Taxonomy
Taxonomy is the science of identifying and naming species, and arranging them into a classification. The field of taxonomy, sometimes referred to as "biological taxonomy", revolves around the description and use of taxonomic units, known as taxa...
, a domain (also superregnum, superkingdom, empire, or regio) is the highest taxonomic rank
Taxonomic rank
In biological classification, rank is the level in a taxonomic hierarchy. Examples of taxonomic ranks are species, genus, family, and class. Each rank subsumes under it a number of less general categories...
of organism
Organism
In biology, an organism is any contiguous living system . In at least some form, all organisms are capable of response to stimuli, reproduction, growth and development, and maintenance of homoeostasis as a stable whole.An organism may either be unicellular or, as in the case of humans, comprise...
s, higher than a kingdom
Kingdom (biology)
In biology, kingdom is a taxonomic rank, which is either the highest rank or in the more recent three-domain system, the rank below domain. Kingdoms are divided into smaller groups called phyla or divisions in botany...
. According to the three-domain system
Three-domain system
The three-domain system is a biological classification introduced by Carl Woese in 1977 that divides cellular life forms into archaea, bacteria, and eukaryote domains. In particular, it emphasizes the separation of prokaryotes into two groups, originally called Eubacteria and Archaebacteria...
of Carl Woese
Carl Woese
Carl Richard Woese is an American microbiologist and physicist. Woese is famous for defining the Archaea in 1977 by phylogenetic taxonomy of 16S ribosomal RNA, a technique pioneered by Woese and which is now standard practice. He was also the originator of the RNA world hypothesis in 1977,...
, introduced in 1990, the Tree of Life
Tree of life (science)
Charles Darwin proposed that phylogeny, the evolutionary relatedness among species through time, was expressible as a metaphor he termed the Tree of Life...
consists of three domains: Archaea
Archaea
The Archaea are a group of single-celled microorganisms. A single individual or species from this domain is called an archaeon...
, Bacteria
Bacteria
Bacteria are a large domain of prokaryotic microorganisms. Typically a few micrometres in length, bacteria have a wide range of shapes, ranging from spheres to rods and spirals...
and Eukarya. The arrangement of taxa
Taxon
|thumb|270px|[[African elephants]] form a widely-accepted taxon, the [[genus]] LoxodontaA taxon is a group of organisms, which a taxonomist adjudges to be a unit. Usually a taxon is given a name and a rank, although neither is a requirement...
reflects the fundamental differences in the genome
Genome
In modern molecular biology and genetics, the genome is the entirety of an organism's hereditary information. It is encoded either in DNA or, for many types of virus, in RNA. The genome includes both the genes and the non-coding sequences of the DNA/RNA....
s.
Alternative classifications of life
Life
Life is a characteristic that distinguishes objects that have signaling and self-sustaining processes from those that do not, either because such functions have ceased , or else because they lack such functions and are classified as inanimate...
so far proposed include:
- The two-empire systemTwo-empire systemThe two-empire system was the top-level biological classification system in general use before the establishment of the three-domain system. It classified life into Prokaryota and Eukaryota...
or superdomain system, with top-level groupings of Prokaryota (or MoneraMoneraMonera is a superseded kingdom that contains unicellular organisms without a nucleus , such as bacteria....
) and EukaryotaEukaryoteA eukaryote is an organism whose cells contain complex structures enclosed within membranes. Eukaryotes may more formally be referred to as the taxon Eukarya or Eukaryota. The defining membrane-bound structure that sets eukaryotic cells apart from prokaryotic cells is the nucleus, or nuclear...
. - The six-kingdom system with top-level groupings of EubacteriaBacteriaBacteria are a large domain of prokaryotic microorganisms. Typically a few micrometres in length, bacteria have a wide range of shapes, ranging from spheres to rods and spirals...
, ArchaebacteriaArchaeaThe Archaea are a group of single-celled microorganisms. A single individual or species from this domain is called an archaeon...
, ProtistProtistProtists are a diverse group of eukaryotic microorganisms. Historically, protists were treated as the kingdom Protista, which includes mostly unicellular organisms that do not fit into the other kingdoms, but this group is contested in modern taxonomy...
a, FungiFungusA fungus is a member of a large group of eukaryotic organisms that includes microorganisms such as yeasts and molds , as well as the more familiar mushrooms. These organisms are classified as a kingdom, Fungi, which is separate from plants, animals, and bacteria...
, PlantPlantPlants are living organisms belonging to the kingdom Plantae. Precise definitions of the kingdom vary, but as the term is used here, plants include familiar organisms such as trees, flowers, herbs, bushes, grasses, vines, ferns, mosses, and green algae. The group is also called green plants or...
ae, and AnimalAnimalAnimals are a major group of multicellular, eukaryotic organisms of the kingdom Animalia or Metazoa. Their body plan eventually becomes fixed as they develop, although some undergo a process of metamorphosis later on in their life. Most animals are motile, meaning they can move spontaneously and...
ia. - The three-empire systemThree-domain systemThe three-domain system is a biological classification introduced by Carl Woese in 1977 that divides cellular life forms into archaea, bacteria, and eukaryote domains. In particular, it emphasizes the separation of prokaryotes into two groups, originally called Eubacteria and Archaebacteria...
(Eubacteria, ArchaeaArchaeaThe Archaea are a group of single-celled microorganisms. A single individual or species from this domain is called an archaeon...
, Eukarya) with five supergroups in the Eukarya (Unikonta, Excavata, Chromalveolata, RhizariaRhizariaThe Rhizaria are a species-rich supergroup of unicellular eukaryotes. This supergroup was proposed by Cavalier-Smith in 2002. They vary considerably in form, but for the most part they are amoeboids with filose, reticulose, or microtubule-supported pseudopods...
and ArchaeplastidaArchaeplastidaThe Archaeplastida are a major group of eukaryotes, comprising the red and green algae and the land plants, together with a small group called the glaucophytes. The plastids of all of these organisms are surrounded by two membranes, suggesting they developed directly from endosymbiotic...
)
None of the three systems currently include non-cellular life
Non-cellular life
Non-cellular life is life that exists without a cellular structure. This term presumes the phylogenetic scientific classification of viruses as lifeforms....
.