Chaetosphaeridium globosum
Encyclopedia
Chaetosphaeridium globosum is a one-celled algae
Algae
Algae are a large and diverse group of simple, typically autotrophic organisms, ranging from unicellular to multicellular forms, such as the giant kelps that grow to 65 meters in length. They are photosynthetic like plants, and "simple" because their tissues are not organized into the many...

 which is thought to represent an ancient lineage of the green plants. This organism exists in a filamentous
Filamentation
Filamentation is the anomalous growth of certain bacteria, such as E. coli, in which cells continue to elongate but do not divide . Bacterial filamentation is often observed as a result of bacteria responding to various stresses, including DNA damage or inhibition of replication...

 form with one flagella per cell
Cell (biology)
The cell is the basic structural and functional unit of all known living organisms. It is the smallest unit of life that is classified as a living thing, and is often called the building block of life. The Alberts text discusses how the "cellular building blocks" move to shape developing embryos....

. It is a freshwater
Freshwater
Fresh water is naturally occurring water on the Earth's surface in ice sheets, ice caps, glaciers, bogs, ponds, lakes, rivers and streams, and underground as groundwater in aquifers and underground streams. Fresh water is generally characterized by having low concentrations of dissolved salts and...

 species. The flagellum is covered in scales in a 3-prong irregular shape called ‘maple leafs’. The cells are usually 11 to 18μm in diameter and with one pyrenoid. Each cell bears long bristle.

Researchers have found that the mitochondrial DNA
Mitochondrial DNA
Mitochondrial DNA is the DNA located in organelles called mitochondria, structures within eukaryotic cells that convert the chemical energy from food into a form that cells can use, adenosine triphosphate...

 of Chaetosphaeridium is markedly different than that of land plants, suggesting that the mitochondria of land plants evolved significantly after the common ancestor between them and living green algae
Green algae
The green algae are the large group of algae from which the embryophytes emerged. As such, they form a paraphyletic group, although the group including both green algae and embryophytes is monophyletic...

. A very slight similarity exists between liverwort
Liverwort
Liverwort may refer to either*Marchantiophyta, a division of non-vascular plants*Hepatica, a genus of spring flowersliverworts are part of the bryophytes group and the bryophytes of the PlantaeIn the bryophytres group their are mosses too....

 mtDNA and Chaetosphaeridium. The chloroplast
Chloroplast
Chloroplasts are organelles found in plant cells and other eukaryotic organisms that conduct photosynthesis. Chloroplasts capture light energy to conserve free energy in the form of ATP and reduce NADP to NADPH through a complex set of processes called photosynthesis.Chloroplasts are green...

 DNA is markedly similar, however, indicating that a close relationship had existed between the Viridiplantae
Viridiplantae
Viridiplantae are a clade comprising the green algae and land plants.In some classification systems they have been treated as a kingdom, under various names, e.g. Viridiplantae, Chlorobionta, or simply Plantae, the latter expanding the traditional Plant Kingdom to include the green algae...

 and the clade
Clade
A clade is a group consisting of a species and all its descendants. In the terms of biological systematics, a clade is a single "branch" on the "tree of life". The idea that such a "natural group" of organisms should be grouped together and given a taxonomic name is central to biological...

 that includes Chaetosphaeridium. This seems to argue that chloroplasts in green plants originated in prehistoric green algae; the family which includes Chaetosphaeridium globosum.

Chloroplasts are known to be captured (symbiotic) cyanobacteria with their own genome
Genome
In modern molecular biology and genetics, the genome is the entirety of an organism's hereditary information. It is encoded either in DNA or, for many types of virus, in RNA. The genome includes both the genes and the non-coding sequences of the DNA/RNA....

. Part of this genome has been transferred to the nucleus and part has been retained in the chloroplast for the continuation of metabolic processes. This symbiosis, now proven by modern genomics
Genomics
Genomics is a discipline in genetics concerning the study of the genomes of organisms. The field includes intensive efforts to determine the entire DNA sequence of organisms and fine-scale genetic mapping efforts. The field also includes studies of intragenomic phenomena such as heterosis,...

, has shown us how Chaetosphaeridium globosum links ancient cyanobacteria with modern green plants like those in our garden.
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