Bakonydraco
Encyclopedia
Bakonydraco is a genus
of azhdarchid
pterosaur
of the Santonian
-age Upper Cretaceous Csehbánya Formation
of the Bakony Mountains, Iharkút, Veszprém
, western Hungary
.
The genus was named in 2005 by David Weishampel, Atilla Ősi and Jianu Coralia. The type species
is Bakonydraco galaczi. The genus name refers to the Bakony Mountains and to Latin
draco, "dragon
". The specific epithet galaczi honors Professor András Galácz, who helped the authors in the Iharkút Research Program, where fossils are since 2000 found in open-pit mining
of bauxite
, among them the remains of pterosaurs, the first ever discovered in Hungary.
Bakonydraco is based on holotype
MTM Gyn/3, a nearly complete mandibula, a fusion of the lower jaws
. Also assigned to it, as paratype
, is MTM Gyn/4, 21: parts from another jaw's symphysis
(the front parts, having fused into a single blade-like structure, of the two lower jaws); azhdarchid wing
bones and neck
vertebrae from the same area may also belong to it.
The lower jaws are toothless and the two halves of the mandibula are frontally fused for about half of its overall length, forming a long, pointed section that is compressed side-to-side and also expanded vertically, giving it a somewhat spear
head- or arrow
head-like shape from the side. This expansion occurs both on the lower edge and on the top surface, where the most extreme point corresponds with a transverse ridge which separates the straight back half of the symphysis from the pointed end in the front. The jaws of MTM Gyn/3 are 29 centimeters long (11.4 inches), and the wingspan
of the genus is estimated to be 3.5 to 4 meters (11.5 to 13.1 feet), which is medium-sized for a pterosaur. Because the jaws are relatively taller than other azhdarchids, and reminiscent of Tapejara, Bakonydraco may have fed differently than other azdarchids. It could have been a piscivore
(feeding on small fish
), or a frugivore
. The discovery of this genus establishes the presence of azhdarchids in the Late Cretaceous of Hungary, suggesting that there as elsewhere they had become the dominant pterosaurs.
Genus
In biology, a genus is a low-level taxonomic rank used in the biological classification of living and fossil organisms, which is an example of definition by genus and differentia...
of azhdarchid
Azhdarchidae
Azhdarchidae is a family of pterosaurs known primarily from the late Cretaceous Period, though an isolated vertebrae apparently from an azhdarchid is known from the early Cretaceous as well...
pterosaur
Pterosaur
Pterosaurs were flying reptiles of the clade or order Pterosauria. They existed from the late Triassic to the end of the Cretaceous Period . Pterosaurs are the earliest vertebrates known to have evolved powered flight...
of the Santonian
Santonian
The Santonian is an age in the geologic timescale or a chronostratigraphic stage. It is a subdivision of the Late Cretaceous epoch or Upper Cretaceous series. It spans the time between 85.8 ± 0.7 mya and 83.5 ± 0.7 mya...
-age Upper Cretaceous Csehbánya Formation
Csehbánya Formation
The Csehbánya Formation is a geological formation in Hungary whose strata date back to the Late Cretaceous. Dinosaur remains are among the fossils that have been recovered from the formation.-Vertebrate paleofauna:...
of the Bakony Mountains, Iharkút, Veszprém
Veszprém (county)
Veszprém is the name of an administrative county in Hungary. Veszprém is also the name of the capital city of Veszprém county.-Veszprém county:...
, western Hungary
Hungary
Hungary , officially the Republic of Hungary , is a landlocked country in Central Europe. It is situated in the Carpathian Basin and is bordered by Slovakia to the north, Ukraine and Romania to the east, Serbia and Croatia to the south, Slovenia to the southwest and Austria to the west. The...
.
The genus was named in 2005 by David Weishampel, Atilla Ősi and Jianu Coralia. The type species
Type species
In biological nomenclature, a type species is both a concept and a practical system which is used in the classification and nomenclature of animals and plants. The value of a "type species" lies in the fact that it makes clear what is meant by a particular genus name. A type species is the species...
is Bakonydraco galaczi. The genus name refers to the Bakony Mountains and to Latin
Latin
Latin is an Italic language originally spoken in Latium and Ancient Rome. It, along with most European languages, is a descendant of the ancient Proto-Indo-European language. Although it is considered a dead language, a number of scholars and members of the Christian clergy speak it fluently, and...
draco, "dragon
Dragon
A dragon is a legendary creature, typically with serpentine or reptilian traits, that feature in the myths of many cultures. There are two distinct cultural traditions of dragons: the European dragon, derived from European folk traditions and ultimately related to Greek and Middle Eastern...
". The specific epithet galaczi honors Professor András Galácz, who helped the authors in the Iharkút Research Program, where fossils are since 2000 found in open-pit mining
Open-pit mining
Open-pit mining or opencast mining refers to a method of extracting rock or minerals from the earth by their removal from an open pit or borrow....
of bauxite
Bauxite
Bauxite is an aluminium ore and is the main source of aluminium. This form of rock consists mostly of the minerals gibbsite Al3, boehmite γ-AlO, and diaspore α-AlO, in a mixture with the two iron oxides goethite and hematite, the clay mineral kaolinite, and small amounts of anatase TiO2...
, among them the remains of pterosaurs, the first ever discovered in Hungary.
Bakonydraco is based on holotype
Holotype
A holotype is a single physical example of an organism, known to have been used when the species was formally described. It is either the single such physical example or one of several such, but explicitly designated as the holotype...
MTM Gyn/3, a nearly complete mandibula, a fusion of the lower jaws
Gnathostomata
Gnathostomata is the group of vertebrates with jaws. The term derives from Greek γνάθος "jaw" + στόμα "mouth". Gnathostome diversity comprises roughly 60,000 species, which accounts for 99% of all living vertebrates...
. Also assigned to it, as paratype
Paratype
Paratype is a technical term used in the scientific naming of species and other taxa of organisms. The exact meaning of the term paratype when it is used in zoology is not the same as the meaning when it is used in botany...
, is MTM Gyn/4, 21: parts from another jaw's symphysis
Symphysis menti
The external surface of the mandible is marked in the median line by a faint ridge, indicating the symphysis menti, mandibular symphysis, or line of junction of the two pieces of which the bone is composed at an early period of life....
(the front parts, having fused into a single blade-like structure, of the two lower jaws); azhdarchid wing
Wing
A wing is an appendage with a surface that produces lift for flight or propulsion through the atmosphere, or through another gaseous or liquid fluid...
bones and neck
Neck
The neck is the part of the body, on many terrestrial or secondarily aquatic vertebrates, that distinguishes the head from the torso or trunk. The adjective signifying "of the neck" is cervical .-Boner anatomy: The cervical spine:The cervical portion of the human spine comprises seven boney...
vertebrae from the same area may also belong to it.
The lower jaws are toothless and the two halves of the mandibula are frontally fused for about half of its overall length, forming a long, pointed section that is compressed side-to-side and also expanded vertically, giving it a somewhat spear
Spear
A spear is a pole weapon consisting of a shaft, usually of wood, with a pointed head.The head may be simply the sharpened end of the shaft itself, as is the case with bamboo spears, or it may be made of a more durable material fastened to the shaft, such as flint, obsidian, iron, steel or...
head- or arrow
Arrow
An arrow is a shafted projectile that is shot with a bow. It predates recorded history and is common to most cultures.An arrow usually consists of a shaft with an arrowhead attached to the front end, with fletchings and a nock at the other.- History:...
head-like shape from the side. This expansion occurs both on the lower edge and on the top surface, where the most extreme point corresponds with a transverse ridge which separates the straight back half of the symphysis from the pointed end in the front. The jaws of MTM Gyn/3 are 29 centimeters long (11.4 inches), and the wingspan
Wingspan
The wingspan of an airplane or a bird, is the distance from one wingtip to the other wingtip. For example, the Boeing 777 has a wingspan of about ; and a Wandering Albatross caught in 1965 had a wingspan of , the official record for a living bird.The term wingspan, more technically extent, is...
of the genus is estimated to be 3.5 to 4 meters (11.5 to 13.1 feet), which is medium-sized for a pterosaur. Because the jaws are relatively taller than other azhdarchids, and reminiscent of Tapejara, Bakonydraco may have fed differently than other azdarchids. It could have been a piscivore
Piscivore
A piscivore is a carnivorous animal which eats primarily fish. Piscivory was the diet of early tetrapods , insectivory came next, then in time reptiles added herbivory....
(feeding on small fish
Fish
Fish are a paraphyletic group of organisms that consist of all gill-bearing aquatic vertebrate animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish, as well as various extinct related groups...
), or a frugivore
Frugivore
A frugivore is a fruit eater. It can be any type of herbivore or omnivore where fruit is a preferred food type. Because approximately 20% of all mammalian herbivores also eat fruit, frugivory is considered to be common among mammals. Since frugivores eat a lot of fruit they are highly dependent...
. The discovery of this genus establishes the presence of azhdarchids in the Late Cretaceous of Hungary, suggesting that there as elsewhere they had become the dominant pterosaurs.