Tilia johnsoni
Encyclopedia
Tilia johnsoni is an extinct species
Species
In biology, a species is one of the basic units of biological classification and a taxonomic rank. A species is often defined as a group of organisms capable of interbreeding and producing fertile offspring. While in many cases this definition is adequate, more precise or differing measures are...

 of linden
Tilia
Tilia is a genus of about 30 species of trees native throughout most of the temperate Northern Hemisphere. The greatest species diversity is found in Asia, and the genus also occurs in Europe and eastern North America, but not western North America...

 tree
Tree
A tree is a perennial woody plant. It is most often defined as a woody plant that has many secondary branches supported clear of the ground on a single main stem or trunk with clear apical dominance. A minimum height specification at maturity is cited by some authors, varying from 3 m to...

 that was native to Washington state, USA and British Columbia
British Columbia
British Columbia is the westernmost of Canada's provinces and is known for its natural beauty, as reflected in its Latin motto, Splendor sine occasu . Its name was chosen by Queen Victoria in 1858...

, Canada. This species lived around 49 million years ago during the Early Eocene
Ypresian
In the geologic timescale the Ypresian is the oldest age or lowest stratigraphic stage of the Eocene. It spans the time between and , is preceded by the Thanetian age and is followed by the Eocene Lutetian age....

. The species was named in honor of Dr. Kirk R. Johnson, paleobotanist who assisted with collecting from the Klondike Mountain Formation
Klondike Mountain Formation
The Klondike Mountain Formation is an early Eocene geologic formation that records the sedimentation in the southern most of a string of highland subtropical/Temperate lakes in Washington state and British Columbia. The formation is best known for exceptionally well preserved plant and insect fossils...

 at the type locality
Type locality (geology)
Type locality , also called type area or type locale, is the where a particular rock type, stratigraphic unit, fossil or mineral species is first identified....

 in Republic, Washington
Republic, Washington
Republic is a city in Ferry County, Washington, United States. The population was 1,073 at the 2010 census, a 12.5% increase over the 2000 Census. It is the county seat of Ferry County.-History:...

. T. johnsoni is the oldest recognized species of the modern Tilia genus in the fossil record to be represented by macroscopic fossils. Fossil pollen assigned to the genus has been recognized from the Palaeocene
Paleocene
The Paleocene or Palaeocene, the "early recent", is a geologic epoch that lasted from about . It is the first epoch of the Palaeogene Period in the modern Cenozoic Era...

 - Eocene
Eocene
The Eocene Epoch, lasting from about 56 to 34 million years ago , is a major division of the geologic timescale and the second epoch of the Paleogene Period in the Cenozoic Era. The Eocene spans the time from the end of the Palaeocene Epoch to the beginning of the Oligocene Epoch. The start of the...

boundary.
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