Rhomboid fossa
Encyclopedia
The anterior part of the fourth ventricle
Fourth ventricle
The fourth ventricle is one of the four connected fluid-filled cavities within the human brain. These cavities, known collectively as the ventricular system, consist of the left and right lateral ventricles, the third ventricle, and the fourth ventricle...

 is named, from its shape, the rhomboid fossa, and its anterior wall, formed by the back of the pons
Pons
The pons is a structure located on the brain stem, named after the Latin word for "bridge" or the 16th-century Italian anatomist and surgeon Costanzo Varolio . It is superior to the medulla oblongata, inferior to the midbrain, and ventral to the cerebellum. In humans and other bipeds this means it...

 and medulla oblongata
Medulla oblongata
The medulla oblongata is the lower half of the brainstem. In discussions of neurology and similar contexts where no ambiguity will result, it is often referred to as simply the medulla...

, constitutes the floor of the fourth ventricle.

It is covered by a thin layer of gray substance continuous with that of the medulla spinalis; superficial to this is a thin lamina of neuroglia which constitutes the ependyma of the ventricle and supports a layer of ciliated epithelium.

Parts

The fossa consists of three parts, superior, intermediate, and inferior.
  • The superior part is triangular in shape and limited laterally by the superior cerebellar peduncle; its apex, directed upward, is continuous with the cerebral aqueduct
    Cerebral aqueduct
    The mesencephalic duct, also known as the aqueductus mesencephali, aqueduct of Sylvius or the cerebral aqueduct, contains cerebrospinal fluid , is within the mesencephalon and connects the third ventricle in the diencephalon to the fourth ventricle in the mesencephalon, which is between the pons...

    ; its base is represented by an imaginary line at the level of the upper ends of the superior foveae.

  • The intermediate part extends from this level to that of the horizontal portions of the taeniae of the ventricle; it is narrow above where it is limited laterally by the middle peduncle, but widens below and is prolonged into the lateral recess
    Lateral recess
    The lateral recess is a projection of the fourth ventricle which extends into the inferior cerebellar peduncle of the brainstem. The lateral aperture, an opening in each extremity of the lateral recess, provides a conduit for cerebrospinal fluid to flow from the brain's ventricular system into the...

    es of the ventricle.

  • The inferior part is triangular, and its downwardly directed apex, named the calamus scriptorius, is continuous with the central canal of the closed part of the medulla oblongata.


The sulcus limitans
Sulcus limitans
In the floor of the fourth ventricle, the sulcus limitans separates the cranial nerve motor nuclei from the sensory nuclei . In the superior part of the rhomboid fossa, it corresponds with the lateral limit of the fossa and presents a bluish-gray area, the locus ceruleus In the floor of the fourth...

 forms the lateral boundary of the medial eminence.

Features

In the superior part of the rhomboid fossa it corresponds with the lateral limit of the fossa and presents a bluish-gray area, the locus cæruleus, which owes its color to an underlying patch of deeply pigmented nerve cells, termed the substantia ferruginea.

At the level of the colliculus facialis the sulcus limitans widens into a flattened depression, the superior fovea, and in the inferior part of the fossa appears as a distinct dimple, the inferior fovea.

Lateral to the foveæ is a rounded elevation named the area acustica, which extends into the lateral recess and there forms a feebly marked swelling, the tuberculum acusticum.

Winding around the inferior peduncle and crossing the area acustica and the medial eminence are a number of white strands, the striæ medullares, which form a portion of the cochlear division of the acoustic nerve and disappear into the median sulcus.

Below the inferior fovea, and between the trigonum hypoglossi and the lower part of the area acustica is a triangular dark field, the ala cinerea, which corresponds to the sensory nucleus of the vagus and glossopharyngeal nerves.

The lower end of the ala cinerea is crossed by a narrow translucent ridge, the funiculus separans, and between this funiculus
Funiculus
The cerebrospinal fibers consist of numerous nerve fibers collected together and enclosed in membranous sheaths. A small bundle of fibers, enclosed in a tubular sheath, is called a funiculus; if the nerve is of small size, it may consist only of a single funiculus; but if large, the funiculi are...

 and the clava
Clava
Clava can refer to:* Mu Boötis, a triple star system in the constellation Boötes.* Clava cairn, a type of Bronze Age circular chamber tomb cairn....

, is a small tongue-shaped area, the area postrema.

On section it is seen that the funiculus separans is formed by a strip of thickened ependyma, and the area postrema by loose, highly vascular, neuroglial tissue containing nerve cells of moderate size.
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