Proardea
Encyclopedia
Proardea is an extinct genus
Genus
In biology, a genus is a low-level taxonomic rank used in the biological classification of living and fossil organisms, which is an example of definition by genus and differentia...

 of heron
Heron
The herons are long-legged freshwater and coastal birds in the family Ardeidae. There are 64 recognised species in this family. Some are called "egrets" or "bitterns" instead of "heron"....

, containing a single species
Species
In biology, a species is one of the basic units of biological classification and a taxonomic rank. A species is often defined as a group of organisms capable of interbreeding and producing fertile offspring. While in many cases this definition is adequate, more precise or differing measures are...

, Proardea amissa ("Lost Proto-heron"). It stood about 70 cm (2 ft 4 in) tall and was very similar to a modern heron
Heron
The herons are long-legged freshwater and coastal birds in the family Ardeidae. There are 64 recognised species in this family. Some are called "egrets" or "bitterns" instead of "heron"....

 in shape. The species is known from rather fragmentary fossil
Fossil
Fossils are the preserved remains or traces of animals , plants, and other organisms from the remote past...

s in the area of Quercy
Quercy
Quercy is a former province of France located in the country's southwest, bounded on the north by Limousin, on the west by Périgord and Agenais, on the south by Gascony and Languedoc, and on the east by Rouergue and Auvergne....

, France
France
The French Republic , The French Republic , The French Republic , (commonly known as France , is a unitary semi-presidential republic in Western Europe with several overseas territories and islands located on other continents and in the Indian, Pacific, and Atlantic oceans. Metropolitan France...

; dated remains are from Pech Desse, a Late Oligocene
Oligocene
The Oligocene is a geologic epoch of the Paleogene Period and extends from about 34 million to 23 million years before the present . As with other older geologic periods, the rock beds that define the period are well identified but the exact dates of the start and end of the period are slightly...

 locality, but the original fossil, a single right tarsometatarsus
Tarsometatarsus
The tarsometatarsus is a bone that is found in the lower leg of certain tetrapods, namely birds.It is formed from the fusion of several bones found in other types of animals, and homologous to the mammalian tarsal and metatarsal bones...

 (MNHN QU-15720), may have come from deposits as early as Late Eocene
Eocene
The Eocene Epoch, lasting from about 56 to 34 million years ago , is a major division of the geologic timescale and the second epoch of the Paleogene Period in the Cenozoic Era. The Eocene spans the time from the end of the Palaeocene Epoch to the beginning of the Oligocene Epoch. The start of the...

 in age.

Proardea was apparently closely related to the true herons and egrets (Ardeinae). As these genera are only known from the Miocene
Miocene
The Miocene is a geological epoch of the Neogene Period and extends from about . The Miocene was named by Sir Charles Lyell. Its name comes from the Greek words and and means "less recent" because it has 18% fewer modern sea invertebrates than the Pliocene. The Miocene follows the Oligocene...

 onwards, Proardea possibly was a direct ancestor of today's herons and/or egrets. However, the Miocene genus Proardeola is closely related, or perhaps even synonymous, with Proardea; the former's single species Proardeola walkeri may thus be Proardea walkeri or even identical with P. amissa. The bird described as Ardea aurelianensis may also be identical with P. amissa, which would in that case become known as Proardeola aurelianensis. Supposed other species of Proardea, P. similis and P. perplexa, are synonyms of the ibis
Ibis
The ibises are a group of long-legged wading birds in the family Threskiornithidae....

 Geronticus perplexus and the phasianid
Phasianidae
The Phasianidae is a family of birds which consists of the pheasants and partridges, including the junglefowl , Old World Quail, francolins, monals and peafowl. The family is a large one, and is occasionally broken up into two subfamilies, the Phasianinae, and the Perdicinae...

 Miogallus altus, respectively.
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