The pileipellis is the uppermost layer of hyphae in the pileus
Pileus (mycology)
The pileus is the technical name for the cap, or cap-like part, of a basidiocarp or ascocarp that supports a spore-bearing surface, the hymenium. The hymenium may consist of lamellae, tubes, or teeth, on the underside of the pileus...

 of a fungal fruit body
In fungi, a basidiocarp, basidiome or basidioma , is the sporocarp of a basidiomycete, the multicellular structure on which the spore-producing hymenium is borne. Basidiocarps are characteristic of the hymenomycetes; rusts and smuts do not produce such structures...

. It covers the trama
Trama (mycology)
In mycology trama is a term for the inner, fleshy portion of a mushroom's basidiocarp, or fruit body. It is distinct from the outer layer of tissue, known as the pileipellis or cuticle, and from the spore-bearing tissue layer known as the hymenium....

, the fleshy tissue of the fruit body. The pileipellis is more or less synonymous with the cuticle, but the cuticle generally describes this layer as a macroscopic feature, while pileipellis refers to this structure as a microscopic layer. Pileipellis type is an important character in the identification of fungi. Pileipellis types include the cutis, trichoderm, epithelium, and hymeniderm types.


A cutis is a type of pileipellis characterized by hyphae that are repent, that is, that run parallel to the pileus surface. In an ixocutis, the hyphae are gelatinous.


In a trichoderm, the outermost hyphae emerge roughly parallel, like hairs, perpendicular to the cap surface. The prefix "tricho-" comes from a Greek word for "hair". In an ixotrichodermium, the outermost hyphae are gelatinous.


An epithelium is a pileipellis consisting of rounded cells in multiple layers, often connected in chains, and sometimes breaking off.


Also called hymeniform, or palisade at times. When viewed from above, a hymeniderm or "cellular cuticle" appears to be paved with roughly circular polygonal elements (similar to the fertile cells of the actual hymenium
The hymenium is the tissue layer on the hymenophore of a fungal fruiting body where the cells develop into basidia or asci, which produce spores. In some species all of the cells of the hymenium develop into basidia or asci, while in others some cells develop into sterile cells called cystidia or...

on the gills). The elements may be globular cells or may be the tips of hyphae extending deeper into the surface.

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