Myelencephalon
Encyclopedia
The myelencephalon is categorized as a secondary vesicle in the development
Morphogenesis
Morphogenesis , is the biological process that causes an organism to develop its shape...

 of the central nervous system
Central nervous system
The central nervous system is the part of the nervous system that integrates the information that it receives from, and coordinates the activity of, all parts of the bodies of bilaterian animals—that is, all multicellular animals except sponges and radially symmetric animals such as jellyfish...

. The prefix "myelen" is derived from Greek for medulla (myelos). The myelencephalon differentiates primarily into the medulla oblongata
Medulla oblongata
The medulla oblongata is the lower half of the brainstem. In discussions of neurology and similar contexts where no ambiguity will result, it is often referred to as simply the medulla...

 and a caudal portion of the fourth ventricle
Fourth ventricle
The fourth ventricle is one of the four connected fluid-filled cavities within the human brain. These cavities, known collectively as the ventricular system, consist of the left and right lateral ventricles, the third ventricle, and the fourth ventricle...

, but will also contain portions of the following cranial nerves: vestibulocochlear nerve
Vestibulocochlear nerve
The vestibulocochlear nerve is the eighth of twelve cranial nerves, and is responsible for transmitting sound and equilibrium information from the inner ear to the brain...

 (CN VIII), glossopharyngeal nerve
Glossopharyngeal nerve
The glossopharyngeal nerve is the ninth of twelve pairs of cranial nerves . It exits the brainstem out from the sides of the upper medulla, just rostral to the vagus nerve...

 (CN IX), vagus nerve
Vagus nerve
The vagus nerve , also called pneumogastric nerve or cranial nerve X, is the tenth of twelve paired cranial nerves...

 (CN X), accessory nerve
Accessory nerve
In anatomy, the accessory nerve is a nerve that controls specific muscles of the shoulder and neck. As part of it was formerly believed to originate in the brain, it is considered a cranial nerve...

 (CN XI), and hypoglossal nerve
Hypoglossal nerve
The hypoglossal nerve is the twelfth cranial nerve , leading to the tongue. The nerve arises from the hypoglossal nucleus and emerges from the medulla oblongata in the preolivary sulcus separating the olive and the pyramid. It then passes through the hypoglossal canal...

 (CN XII).

The myelencephalon develops from the primary vesicular structure called the rhombencephalon
Rhombencephalon
The rhombencephalon is a developmental categorization of portions of the central nervous system in vertebrates.The rhombencephalon can be subdivided in a variable number of transversal swellings called rhombomeres...

 (or hindbrain), which is present in all vertebrate embryos. The rhombencephalon normally begins its differentiation into the myelencephalon and the metencephalon
Metencephalon
The metencephalon is a developmental categorization of portions of the central nervous system. The metencephalon is composed of the pons and the cerebellum; contains a portion of the fourth ventricle; and the trigeminal nerve , abducens nerve , facial nerve , and a portion of the vestibulocochlear...

 at approximately 5 weeks of gestational development in humans. Evolutionarily, the myelencephalon is the area of the brain which is the most ancestral, eventually controlling visceral mechanisms (basic bodily functions) such as breathing, heart and blood vessel activity, digestions, and peristalsis
Peristalsis
Peristalsis is a radially symmetrical contraction and relaxation of muscles which propagates in a wave down the muscular tube, in an anterograde fashion. In humans, peristalsis is found in the contraction of smooth muscles to propel contents through the digestive tract. Earthworms use a similar...

.
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