Kambara
Encyclopedia
Kambara is an extinct genus
Genus
In biology, a genus is a low-level taxonomic rank used in the biological classification of living and fossil organisms, which is an example of definition by genus and differentia...

 of mekosuchine crocodylian that lived during the Eocene
Eocene
The Eocene Epoch, lasting from about 56 to 34 million years ago , is a major division of the geologic timescale and the second epoch of the Paleogene Period in the Cenozoic Era. The Eocene spans the time from the end of the Palaeocene Epoch to the beginning of the Oligocene Epoch. The start of the...

 and Oligocene
Oligocene
The Oligocene is a geologic epoch of the Paleogene Period and extends from about 34 million to 23 million years before the present . As with other older geologic periods, the rock beds that define the period are well identified but the exact dates of the start and end of the period are slightly...

 epochs in Australia
Australia
Australia , officially the Commonwealth of Australia, is a country in the Southern Hemisphere comprising the mainland of the Australian continent, the island of Tasmania, and numerous smaller islands in the Indian and Pacific Oceans. It is the world's sixth-largest country by total area...

.

At around 55 million years old, remains of Kambara are among the oldest Tertiary
Tertiary
The Tertiary is a deprecated term for a geologic period 65 million to 2.6 million years ago. The Tertiary covered the time span between the superseded Secondary period and the Quaternary...

 fossils found in Australia (although there are some recent Cretaceous
Cretaceous
The Cretaceous , derived from the Latin "creta" , usually abbreviated K for its German translation Kreide , is a geologic period and system from circa to million years ago. In the geologic timescale, the Cretaceous follows the Jurassic period and is followed by the Paleogene period of the...

 fossils that are twice that age). Kambara is the oldest known mekosuchine. The genus name comes from an Aboriginal term meaning "crocodile".

There are currently four species of Kambara described: the type species
Type species
In biological nomenclature, a type species is both a concept and a practical system which is used in the classification and nomenclature of animals and plants. The value of a "type species" lies in the fact that it makes clear what is meant by a particular genus name. A type species is the species...

 K. murgonensis (Willis & Molnar, 1993), K. implexidens (Salisbury & Willis, 1996), K. molnari (Holt et al., 2005), and K. taraina (Buchanan, 2009). All four species have a generalised crocodylian body plan, growing to sizes similar to the modern Saltwater Crocodile
Saltwater Crocodile
The saltwater crocodile, also known as estuarine or Indo-Pacific crocodile, is the largest of all living reptiles...

, Crocodylus porosus. Kambara shows an interesting characteristic of having multiple bite patterns within the same genus. Kambara murgonensis has a near complete overbite
Malocclusion
A malocclusion is a misalignment of teeth or incorrect relation between the teeth of the two dental arches. The term was coined by Edward Angle, the "father of modern orthodontics", as a derivative of occlusion, which refers to the manner in which opposing teeth meet.-Presentation:Most people have...

, K. implexidens a more interlocking dentition and K. molnarai an intermediate condition. While initially thought to be the most primitive member of an Australasia
Australasia
Australasia is a region of Oceania comprising Australia, New Zealand, the island of New Guinea, and neighbouring islands in the Pacific Ocean. The term was coined by Charles de Brosses in Histoire des navigations aux terres australes...

n radiation of mekosuchine crocodylians, recent studies (Holt, et al., 2007) have suggested that this may not be the case, and that there are at least two separate lineages in Australia.

Fossils have been found at the Murgon fossil site
Murgon fossil site
The Murgon fossil site is a paleontological site of early Eocene age in south-eastern Queensland, Australia. It lies near the town of Murgon, some 270 km north-west of Brisbane...

 in south-eastern Queensland
Queensland
Queensland is a state of Australia, occupying the north-eastern section of the mainland continent. It is bordered by the Northern Territory, South Australia and New South Wales to the west, south-west and south respectively. To the east, Queensland is bordered by the Coral Sea and Pacific Ocean...

.
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