Hypsibema
Encyclopedia
Hypsibema is a little-known genus
Genus
In biology, a genus is a low-level taxonomic rank used in the biological classification of living and fossil organisms, which is an example of definition by genus and differentia...

 of dinosaur
Dinosaur
Dinosaurs are a diverse group of animals of the clade and superorder Dinosauria. They were the dominant terrestrial vertebrates for over 160 million years, from the late Triassic period until the end of the Cretaceous , when the Cretaceous–Paleogene extinction event led to the extinction of...

 from the Late Cretaceous
Late Cretaceous
The Late Cretaceous is the younger of two epochs into which the Cretaceous period is divided in the geologic timescale. Rock strata from this epoch form the Upper Cretaceous series...

 (Campanian
Campanian
The Campanian is, in the ICS' geologic timescale, the fifth of six ages of the Late Cretaceous epoch . The Campanian spans the time from 83.5 ± 0.7 Ma to 70.6 ± 0.6 Ma ...

 stage
Faunal stage
In chronostratigraphy, a stage is a succession of rock strata laid down in a single age on the geologic timescale, which usually represents millions of years of deposition. A given stage of rock and the corresponding age of time will by convention have the same name, and the same boundaries.Rock...

, around 75 million years ago). Its giant fossils were found in the U.S. states of North Carolina
North Carolina
North Carolina is a state located in the southeastern United States. The state borders South Carolina and Georgia to the south, Tennessee to the west and Virginia to the north. North Carolina contains 100 counties. Its capital is Raleigh, and its largest city is Charlotte...

 and Missouri
Missouri
Missouri is a US state located in the Midwestern United States, bordered by Iowa, Illinois, Kentucky, Tennessee, Arkansas, Oklahoma, Kansas and Nebraska. With a 2010 population of 5,988,927, Missouri is the 18th most populous state in the nation and the fifth most populous in the Midwest. It...

. It is believed to be a hadrosaur, although the Missouri remains were first thought to belong to a small sauropod ("Neosaurus", renamed Parrosaurus).
The type species
Type species
In biological nomenclature, a type species is both a concept and a practical system which is used in the classification and nomenclature of animals and plants. The value of a "type species" lies in the fact that it makes clear what is meant by a particular genus name. A type species is the species...

, Hypsibema crassicauda, was described by Edward Drinker Cope
Edward Drinker Cope
Edward Drinker Cope was an American paleontologist and comparative anatomist, as well as a noted herpetologist and ichthyologist. Born to a wealthy Quaker family, Cope distinguished himself as a child prodigy interested in science; he published his first scientific paper at the age of nineteen...

 in 1869. The generic name is derived from Greek υψι/hypsi, "high", and βεμα/bema, "step", as Cope believed that the species walked particularly erect on its toes. The specific name means "with a fat tail" in Latin
Latin
Latin is an Italic language originally spoken in Latium and Ancient Rome. It, along with most European languages, is a descendant of the ancient Proto-Indo-European language. Although it is considered a dead language, a number of scholars and members of the Christian clergy speak it fluently, and...

. The holotype
Holotype
A holotype is a single physical example of an organism, known to have been used when the species was formally described. It is either the single such physical example or one of several such, but explicitly designated as the holotype...

, USNM 7189, consists of a caudal vertebra, a humerus, a tibia and a metatarsal, found in 1869 by North Carolina state geologist professor Washington Carruthers Kerr in the King James marl pits. A second vertebra referred to the species, USNM 6136, was later discovered by Edward Wilber Berry.

A second species, H. missouriensis
Hypsibema missouriensis
Hypsibema missouriensis is a species of plant-eating dinosaur in the genus Hypsibema, and the state dinosaur of the U.S. state Missouri. One of the few official state dinosaurs, bones of the species were discovered in 1942, at what later became known as the Chronister Dinosaur Site near Glen Allen,...

, named by Donald Baird and Jack Horner in 1979 (previously Parrosaurus missouriensis Gilmore 1945), is since 2004 the official state dinosaur of Missouri
Missouri
Missouri is a US state located in the Midwestern United States, bordered by Iowa, Illinois, Kentucky, Tennessee, Arkansas, Oklahoma, Kansas and Nebraska. With a 2010 population of 5,988,927, Missouri is the 18th most populous state in the nation and the fifth most populous in the Midwest. It...

. Its holotype is USNM 16735, a set of vertebrae. Both species are based on fragmentary material, and are today considered dubious
Nomen dubium
In zoological nomenclature, a nomen dubium is a scientific name that is of unknown or doubtful application...

.
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