Hydrotropism
Encyclopedia
Hydrotropism is a plant's growth response in which the direction of growth is determined by a stimulus or gradient in water concentration. A common example is plant roots growing in humid air bending toward a higher relative humidity level. This is of biological significance as it helps to increase efficiency of the plant in its ecosystem. The process of hydrotropism is started by the root cap sensing water and sending a signal to the elongating part of the root. Hydrotropism is difficult to observe in underground roots, since the roots are not readily observable, and root gravitropism
Gravitropism
Gravitropism is a turning or growth movement by a plant or fungus in response to gravity. Charles Darwin was one of the first to scientifically document that roots show positive gravitropism and stems show negative gravitropism. That is, roots grow in the direction of gravitational pull and stems...

 is usually more influential than root hydrotropism. Also, water is not as strongly directional factor as gravity for gravitropism, light for phototropism
Phototropism
Phototropism is directional growth in which the direction of growth is determined by the direction of the light source. In other words, it is the growth and response to a light stimulus. Phototropism is most often observed in plants, but can also occur in other organisms such as fungi...

 and touch for thigmotropism
Thigmotropism
Thigmotropism is a movement in which an organism moves or grows in response to touch or contact stimuli. The prefix thigmo- θιγμος comes from the Greek for "touch". Usually thigmotropism occurs when plants grow around a surface, such as a wall, pot, or trellis. Climbing plants, such as vines,...

. Water readily moves in soil and soil water content is constantly changing so any gradients in soil moisture are not stable.

Thus, root hydrotropism research has mainly been a laboratory phenomenon for roots grown in humid air rather than soil. Its ecological significance in soil-grown roots is unclear because so little hydrotropism research has examined soil-grown roots. Recent identification of a mutant plant that lacks a hydrotropic response may help to elucidate its role in nature. Hydrotropism may have importance for plants grown in space, where it may allow roots to orient themselves in a microgravity environment.

Misconceptions

  • The greater growth of roots in moist soil zones than in dry soil zones is not usually a result of hydrotropism. Hydrotropism requires a root to bend from a drier to a wetter soil zone. Roots require water to grow so roots that happen to be in moist soil will grow and branch much more than those in dry soil.
  • Roots cannot sense water inside intact pipes via hydrotropism and break the pipes to obtain the water.
  • Roots cannot sense water several feet away via hydrotropism and grow toward it. At best hydrotropism probably operates over distances of a couple millimeters.
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