Heteronomous annulation
Encyclopedia
Heteronomous annulation is a characteristic of some arthropod
Arthropod
An arthropod is an invertebrate animal having an exoskeleton , a segmented body, and jointed appendages. Arthropods are members of the phylum Arthropoda , and include the insects, arachnids, crustaceans, and others...

s. It means that segment
Segmentation (biology)
Segmentation in biology refers to either a type of gastrointestinal motility or the division of some animal and plant body plans into a series of repetitive segments. This article will focus on the segmentation of animal body plans, specifically using the examples of the phyla Arthropoda,...

s are differentiated from one another, each having different form so as to serve a different function - for instance, using some limbs for walking and others for feeding.

The trait is first observed in the armoured lobopods. It is thought to be the precursor to segmentation in arthropods, and perhaps resulted from the need to produce serially repeated defensive structures (spines). It can be studied using homeotic genes.
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