Gomphotaria
Encyclopedia
Gomphotaria pugnax was a species of very large shellfish-eating dusignathine walrus
Walrus
The walrus is a large flippered marine mammal with a discontinuous circumpolar distribution in the Arctic Ocean and sub-Arctic seas of the Northern Hemisphere. The walrus is the only living species in the Odobenidae family and Odobenus genus. It is subdivided into three subspecies: the Atlantic...

 found along the coast of what is now California
California
California is a state located on the West Coast of the United States. It is by far the most populous U.S. state, and the third-largest by land area...

, during the late Miocene
Miocene
The Miocene is a geological epoch of the Neogene Period and extends from about . The Miocene was named by Sir Charles Lyell. Its name comes from the Greek words and and means "less recent" because it has 18% fewer modern sea invertebrates than the Pliocene. The Miocene follows the Oligocene...

. It had four tusks, with one pair in the lower and upper jaws, and according to wear on the tusks, G. pugnax hammered shellfish open, rather than simply suck them out of their shells as do modern walruses. In terms of the postcranial skeleton, Gomphotaria and other dusignathine walruses were built more like sea lions than modern walrus. Gomphotaria is a prime example of the extreme diversity that walruses once exhibited.
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