Gegepterus
Encyclopedia
Gegepterus was a genus
Genus
In biology, a genus is a low-level taxonomic rank used in the biological classification of living and fossil organisms, which is an example of definition by genus and differentia...

 of ctenochasmatid
Ctenochasmatidae
Ctenochasmatidae is a group of pterosaurs within the suborder Pterodactyloidea.-Classification:Listing of subfamilies and genera after Unwin 2006 unless otherwise noted.* Family Ctenochasmatidae** Elanodactylus** Gegepterus...

 pterosaur
Pterosaur
Pterosaurs were flying reptiles of the clade or order Pterosauria. They existed from the late Triassic to the end of the Cretaceous Period . Pterosaurs are the earliest vertebrates known to have evolved powered flight...

 from the Early Cretaceous
Early Cretaceous
The Early Cretaceous or the Lower Cretaceous , is the earlier or lower of the two major divisions of the Cretaceous...

-age Yixian Formation
Yixian Formation
The Yixian Formation is a geological formation in Jinzhou, Liaoning, People's Republic of China, that spans 11 million years during the early Cretaceous period...

 of Liaoning
Liaoning
' is a province of the People's Republic of China, located in the northeast of the country. Its one-character abbreviation is "辽" , a name taken from the Liao River that flows through the province. "Níng" means "peace"...

, China
People's Republic of China
China , officially the People's Republic of China , is the most populous country in the world, with over 1.3 billion citizens. Located in East Asia, the country covers approximately 9.6 million square kilometres...

.

The genus was named in 2007 by Wang Xiaolin, Alexander Kellner
Alexander Kellner
Alexander Wilhelm Armin Kellner is a Liechtensteinian/Brazilian paleontologist, a leading expert in the field of the study of pterosaurs....

, Zhou Zhonge and Diogenes de Almeida Campos. The type species
Type species
In biological nomenclature, a type species is both a concept and a practical system which is used in the classification and nomenclature of animals and plants. The value of a "type species" lies in the fact that it makes clear what is meant by a particular genus name. A type species is the species...

 is Gegepterus changi. The genus name is derived from Manchu
Manchu language
Manchu is a Tungusic endangered language spoken in Northeast China; it used to be the language of the Manchu, though now most Manchus speak Mandarin Chinese and there are fewer than 70 native speakers of Manchu out of a total of nearly 10 million ethnic Manchus...

 ge ge, the title of a princess, in reference to the dainty gracility of the specimen, and a Latinised Greek pteron, "wing". The specific name honours female paleontologist Chang Meemann, who over the years established a cordial relationship between the Chinese and Brazilian authors. In 2008 Wang emended the epithet to changae, but such changes are no longer allowed by the ICZN
International Code of Zoological Nomenclature
The International Code of Zoological Nomenclature is a widely accepted convention in zoology that rules the formal scientific naming of organisms treated as animals...

.

It is known from two specimens. The first is holotype
Holotype
A holotype is a single physical example of an organism, known to have been used when the species was formally described. It is either the single such physical example or one of several such, but explicitly designated as the holotype...

 IVPP V 11981, which was in 2001 found in grey shale
Shale
Shale is a fine-grained, clastic sedimentary rock composed of mud that is a mix of flakes of clay minerals and tiny fragments of other minerals, especially quartz and calcite. The ratio of clay to other minerals is variable. Shale is characterized by breaks along thin laminae or parallel layering...

s from the lower part of the formation
Rock formation
This is a list of rock formations that include isolated, scenic, or spectacular surface rock outcrops. These formations are usually the result of weathering and erosion sculpting the existing rock...

 (estimated at 125 million years old), near the city of Beipiao
Beipiao
Beipiao is a city in Chaoyang prefecture, Liaoning province, in Northeast China. It has a population of 202,807. The main industry in the area is coal mining. With vertical shafts of almost 1000m, these are some of the deepest coal mines in China. The coal produced is used for coking...

. It consists of a crushed and damaged partial skeleton of a subadult
Juvenile (organism)
A juvenile is an individual organism that has not yet reached its adult form, sexual maturity or size. Juveniles sometimes look very different from the adult form, particularly in terms of their colour...

 including skull, lower jaws, cervical and sacral vertebrae, ribs, gastralia
Gastralium
Gastralia are dermal bones found in the ventral body wall of crocodilian and Sphenodon species. They are found between the sternum and pelvis, and do not articulate with the vertebrae...

 ("belly ribs"), shoulder girdle and hindlimb remains, along with dark soft tissue remains near the skull and gastralia and in the orbit; unfortunately, the soft tissue remains show no structure except for some small, unbranched fibers at the back of the head. The jaws are very elongated; the snout is flat and concave on top, with a low and thin crest. The forehead slightly projects to the front. The cervicals are elongated.

The authors assigned it to the Ctenochasmatidae on the basis of its long rostrum
Beak
The beak, bill or rostrum is an external anatomical structure of birds which is used for eating and for grooming, manipulating objects, killing prey, fighting, probing for food, courtship and feeding young...

 and numerous needle-like teeth, about 150 in total. This is the first uncontroversial report of the Ctenochasmatidae from the Yixian Formation, as the fossils of other assumed ctenochasmatids have not preserved the dentition. It was at first suspected to be the juvenile of some known species.

In 2011 a second, smaller specimen was described, IVPP V 11972, which increased the known skeletal elements and showed a more extensive covering of hair-like structures.
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