Cystopteris hybrid complex
Encyclopedia

North American Cystopteris Hybrids

The diploid parent species within the genus include:
  • C. bulbifera
    Cystopteris bulbifera
    Cystopteris bulbifera ' is a low-growing rock fern with creeping stems and narrow elongate deltate fronds which grow to 75 cm , native to eastern North America with two disjunct populations in the west. It is found only on calcareous substrates such as limestone. It commonly festoons limestone cave...

  • C. "hemifragilis" (an undiscovered or extinct diploid)
  • C. protrusa
    Cystopteris protrusa
    Cystopteris protrusa is a common fern of eastern North America. Throughout much of its range, it is the most common Cystopteris. It is commonly known as the lowland brittle fern or lowland fragile fern...

  • C. reevesiana


The hybrid species include:
  • C. fragilis
    Cystopteris fragilis
    Cystopteris fragilis is a species of fern known by the common names brittle bladderfern and common fragile fern. It can be found worldwide, generally in shady, moist areas. The leaves are up to 30 or 40 centimeters long and are borne on fleshy petioles. Each leaf is divided into many pairs of...

    (C. "hemifragilis" X C. reevesiana; allotetraploid)
  • C. laurentiana (C. bulbifera X C. fragilis; allohexaploid)
  • C. tennesseensis (C. bulbifera X C. protrusa; allotetraploid)
  • C. tenuis
    Cystopteris tenuis
    Cystopteris tenuis is sometimes known as Mackay's bladder fern or Mackay's fragile fern. It was long considered to be a part of the superspecies for fragile ferns, as Cystopteris fragilis Bernh. var. mackayi Lawson....

    (C. "hemifragilis" X C. protrusa; allotetraploid)
  • C. utahensis (C. reevesiana X C. bulbifera; allotetraploid)


Non-species hybrids include:
  • Cystopteris Xillinoensis (C. bulbifera X C. tenuis; allotriploid)
  • Cystopteris Xwagneri (C. tennesseensis X C. tenuis; allo-allotetraploid)
  • Cystopteris fragilis X Cystopteris tenuis (allo-allotetraploid)
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