Chenanisuchus
Encyclopedia
Chenanisuchus is a genus
Genus
In biology, a genus is a low-level taxonomic rank used in the biological classification of living and fossil organisms, which is an example of definition by genus and differentia...

 of dyrosaurid
Dyrosauridae
Dyrosauridae is a family of extinct neosuchian crocodyliforms that lived from the Late Cretaceous to the Eocene. Fossils of this group have been found in almost every continent, specifically Africa, Asia, Europe, North America and South America....

 crocodyliform
Crocodyliformes
Crocodyliformes is a clade of crurotarsan archosaurs, the group often traditionally referred to as "crocodilians."In 1988, Michael J. Benton and James M. Clark argued that all traditional names for well-known groups of animals should be restricted to their crown clades, that is, used only for...

 from the Late Cretaceous
Late Cretaceous
The Late Cretaceous is the younger of two epochs into which the Cretaceous period is divided in the geologic timescale. Rock strata from this epoch form the Upper Cretaceous series...

 of Mali
Mali
Mali , officially the Republic of Mali , is a landlocked country in Western Africa. Mali borders Algeria on the north, Niger on the east, Burkina Faso and the Côte d'Ivoire on the south, Guinea on the south-west, and Senegal and Mauritania on the west. Its size is just over 1,240,000 km² with...

 and the Late Palaeocene of Sidi Chenane in Morocco
Morocco
Morocco , officially the Kingdom of Morocco , is a country located in North Africa. It has a population of more than 32 million and an area of 710,850 km², and also primarily administers the disputed region of the Western Sahara...

. It was described in 2005, after expeditions uncovered it in 2000.

The type species is C. lateroculi ("lateralis", lateral; "oculi", eyes), in reference to the laterally facing eyes.

Currently, Chenanisuchus is the most basal known dyrosaurid.

Material

Two specimens of C. lateroculi – OCP DEK-GE 262 (holotype
Holotype
A holotype is a single physical example of an organism, known to have been used when the species was formally described. It is either the single such physical example or one of several such, but explicitly designated as the holotype...

, nearly complete skull with mandibular fragments) and OCP DEK-GE 61 (nearly complete skull) – come from the Sidi Chenane area in Morocco, which is Late Palaeocene (Thanetian) in age. Fossils of Chenanisuchus were also found in Maastrichtian
Maastrichtian
The Maastrichtian is, in the ICS' geologic timescale, the latest age or upper stage of the Late Cretaceous epoch or Upper Cretaceous series, the Cretaceous period or system, and of the Mesozoic era or erathem. It spanned from 70.6 ± 0.6 Ma to 65.5 ± 0.3 Ma...

 age strata in Mali
Mali
Mali , officially the Republic of Mali , is a landlocked country in Western Africa. Mali borders Algeria on the north, Niger on the east, Burkina Faso and the Côte d'Ivoire on the south, Guinea on the south-west, and Senegal and Mauritania on the west. Its size is just over 1,240,000 km² with...

, what shows that Chenanisuchus survived the Cretaceous–Tertiary extinction event.

Systematics

Chenanisuchus lateroculi is referred to Dyrosauridae by Jouve et al. (2005), based on three morphological characters:
  • Presence of occipital tuberosities
  • Presence of an anterolateral postorbital process
  • Large participation of the quadratojugal and surangular to the jaw joint

Palaeobiology

Chenanisuchus lateroculi has an estimated adult length between 4 and 4.5 meters, based on the 60 centimeter long skull. It has the shortest snout relative to the dorsal skull length among all dyrosaurids.
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