Antioch Dunes Evening Primrose
Encyclopedia
Oenothera deltoides subsp. howellii, the Antioch Dunes Evening Primrose, is an endangered
Endangered species
An endangered species is a population of organisms which is at risk of becoming extinct because it is either few in numbers, or threatened by changing environmental or predation parameters...

 subspecies of plant in the family Onagraceae
Onagraceae
Onagraceae, also known as the Willowherb family or Evening Primrose family, are a family of flowering plants. The family includes about 640-650 species of herbs, shrubs, and trees in 20-24 genera...

, genus Oenothera
Oenothera
Oenothera is a Genus of about 125 species of annual, biennial and perennial herbaceous flowering plants, native to North and South America. It is the type genus of the family Onagraceae, the Evening Primrose Family. Common names include evening-primrose, suncups, and sundrops.The species vary in...

, and species Oenothera deltoides
Oenothera deltoides
Oenothera deltoides is a species of evening primrose known by several common names, including birdcage evening primrose, basket evening primrose, lion in a cage, and devil's lantern. It is native to the Southwestern United States and northern Mexico, where it grows in sandy habitats from desert to...

.

Description

Oenothera deltoides subsp. howellii is a short-lived perennial plant with thick stems varying from 4 to 40 inches in length. It is multi-branching in form and tends to grow in large tufts. The grayish colored leaves are shaped like a lance
Lance
A Lance is a pole weapon or spear designed to be used by a mounted warrior. The lance is longer, stout and heavier than an infantry spear, and unsuited for throwing, or for rapid thrusting. Lances did not have tips designed to intentionally break off or bend, unlike many throwing weapons of the...

 and are about 1 to 5 inches long with many hairs, primarily short with only a few being longer. Its flowers are white with inch-long petals and yellow stamens. It blooms from March to September. The subspecies is often confused with Oenothera deltoides ssp. cognata that grows in an area southeast of the protected area.

The Antioch Dunes Evening Primrose blooms for only one night. The flowers are a major host plant for a rare species of sweat bee, Sphecodogastra antiochensis.

Distribution and habitat

Oenothera deltoides subsp. howellii is a perennial plant that grows only in an environmentally sensitive sand dune habitat located near the meeting of the Sacramento River
Sacramento River
The Sacramento River is an important watercourse of Northern and Central California in the United States. The largest river in California, it rises on the eastern slopes of the Klamath Mountains, and after a journey south of over , empties into Suisun Bay, an arm of the San Francisco Bay, and...

 and the San Joaquin River
San Joaquin River
The San Joaquin River is the largest river of Central California in the United States. At over long, the river starts in the high Sierra Nevada, and flows through a rich agricultural region known as the San Joaquin Valley before reaching Suisun Bay, San Francisco Bay, and the Pacific Ocean...

 in California
California
California is a state located on the West Coast of the United States. It is by far the most populous U.S. state, and the third-largest by land area...

.

In 1979 Oenothera deltoides subsp. howellii, the Antioch Dunes Evening Primrose, was declared an endangered species. Currently, its habitat is restricted to the Antioch Dunes National Wildlife Refuge
Antioch Dunes National Wildlife Refuge
Antioch Dunes National Wildlife Refuge is a sensitive sand dune habitat located near the city of Antioch, California on the south shore of the San Joaquin River. It serves as a refuge for three endangered species of plants and insects, and is closed to the public except for tours and events...

, the last remnant of an isolated nine kilometer stretch of sand dune habitat. The refuge was formed in 1980 to prevent extinction of this plant, as well as Lange's metalmark butterfly (Apodemia mormo langei) and the Contra Costa wallflower (Erysium capitatum var. angustatum). This refuge is the first national wildlife refuge in the U.S. formed to protect endangered plants and insects, and it is open for viewing to the public only under supervision because of the sensitive status of these species.

Endangered status

In 2007 a five-year review of its endangered status was completed, and its status was continued unchanged.
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